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. 2021 Sep 23;11:18942. doi: 10.1038/s41598-021-97600-0

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© The Author(s) 2021

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Timetree of the five species of Danionella illustrating relationships of D. cerebrum (left), differences in external appearance of preserved specimens (middle), and sexual dimorphisms in the skeleton of the Weberian apparatus (right, double column) in cleared and double stained specimens. Preserved specimens (middle) from top: Danionella dracula, BMNH 2008.1.1.1, male, holotype, BMNH.1.1.2–99, female, paratype, D. priapus, BMNH 2009.9.9.1, male, holotype, BMNH 2009.9.9.2–37, paratype, female; D. translucida NRM 32235, male and female paratypes; D. mirifica, USNM 372848, male and female paratypes; D. cerebrum, MTD 39985, male, paratype, BMNH 2021.8.30.1, female, holotype. Cleared and stained specimens (scale bar 0.1 mm), males, left column from top: D. dracula, BMNH 2008.1.1.100–119, 16.2 mm; D. priapus, BMNH 2009.9.9.38–43, 16.5 mm; D. translucida, MTD 39992, 9.8 mm; D. mirifica, USNM 372848, 13.2 mm; D. cerebrum, MTD 39986, 11.7 mm; black arrowheads mark connection between lateral process and outer arm of os suspensorium, black stars mark connecting flanges between inner and outer arms of os suspensorium, and red arrows marks posterior extension of inner arm of os suspensorium covering swimbladder dorsally. Females, right column from top: D. dracula, BMNH 2008.1.1.100–119, 14.7 mm; D. priapus, BMNH 2009.9.9.38–43, 14.8 mm; D. translucida, MTD 39992, 11.2 mm; D. mirifica, USNM 372848, 13.2 mm, D. cerebrum, MTD 39986, 11.7 mm.