(A) Diagram of the ELMOD_C (At1g67400), ELMOD_D (At3g43400), ELMOD_E (At1g03620), and ELMOD_F (At3g03610) genes. T-DNA insertion sites are indicated for each gene. For ELMOD_E, CRISPR alleles (elmod-eCR) were also generated in the wild-type and mcr backgrounds. Both alleles had frame shift mutations. The 20 bp target sequence next to the underlined protospacer adjacent motif is indicated in bold. Nucleotide and amino acid changes are indicated with red capital letters. (B–F’) Pollen grains of single T-DNA insertion mutants of ELMOD_C, ELMOD_D, ELMOD_E, ELMOD_F, and the CRISPR/Cas9 mutant of ELMOD_E (elmod_eCR). (G–J’) Pollen grains of the double mutants mcr elmod_eCR (G, G’), mcr elmod_c (H, H’), mcr elmod_d (I, I’), and mcr elmod_f (J, J’). Apertures are indicated with arrowheads. Scale bars, 10 μm.