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. 2021 Oct 1;9:e12160. doi: 10.7717/peerj.12160

Table 8. Qualitative comparisons among adult humeri.

Taxon Cross-sectional shape Presence of ICL and OCL Composition of middle cortical layer Relative vascular density Primary vascular orientation
California Quail1
(Callipepla californica)
circular to ovate yes; OCL thicker (~1/3 of cortex) but ICL more distinct woven/incipient fibrolamellar moderate longitudinal
Wild Turkey1
(Meleagris gallopavo)
rounded triangle yes; ICL thicker and more distinct fibrolamellar high circumferential/
plexiform
Western Gull2
(Larus occidentalis)
circular yes; ICL thicker and more distinct fibrolamellar high reticular
American Kestrel3
(Falco sparverius)
circular yes; equal thickness, ICL more distinct woven w/simple canals and regional fibrolamellar moderate longitudinal/
reticular
Red-tailed Hawk3
(Buteo jamaicensis)
circular yes; ICL thicker and more distinct fibrolamellar high longitudinal
White-tailed Kite3
(Elanus leucurus)
circular yes; OCL thicker, ICL more distinct fibrolamellar bone with regional woven bone w/simple canals moderate longitudinal
Barn Owl3 (Tyto alba) circular yes; equal thickness, ICL more distinct fibrolamellar high reticular/
circumferential
Great-horned Owl3
(Bubo virginianus)
circular yes; ICL thicker and more distinct fibrolamellar high longitudinal
Mourning Dove4
(Zenaida macroura)
ovate yes; ICL incipient, OCL 1/3-1/2 cortex weakly woven with primary osteons (middle layer is of equal thickness to OCL in some areas) moderate longitudinal
Anna’s Hummingbird5
(Calypte anna)
ovate (main medullary cavity) yes; OCL dominates cortex none avascular
Green-cheeked conure5
(Pyrrhura molinae)
circular yes (but ICL thin/irregular); OCL thicker woven bone with primary vascular canals; patches of fibrolamellar moderate-low longitudinal
House Finch5
(Haemorhous mexicanus)
circular yes; OCL is thicker, ~1/3-1/2 of cortex woven bone with primary vascular canals; patches of fibrolamellar moderate-low longitudinal

Note:

Vascular density in adults was ubiquitously lower than in younger growth stages, thus relative vascular density among adults only is reported here. “Middle cortical layer” refers to the portion of the cortex located between the OCL and the ICL. Descriptions of the ICL as “more distinct” refer to instances in which the ICL is more advanced in development than the OCL, and to the very high level of organization of tissue in this layer (which appears strongly lamellated and has a distinct, high birefringence). Developmental mode is indicated by color number superscript: orange1, precocial; yellow2, semi-precocial; green3, semi-altricial 1; blue4, semi-altricial 2; purple5, altricial.