(A) Hamp2CreER Tomato labeling at 1, 26, and 52 weeks. Endogenous GS immunofluorescence shown in green marks the CV region. Cartoon depicting the pattern of Tomato labeling in Hamp2-CreER livers over time (bottom right).(B) Manual measurements of Hamp2-CreER–labeled Tomatopositive cells as a function of position along the CV-to-PV axis. (C) The percent area labeled by Hamp2-CreER at 1, 26, and 52 weeks after tamoxifen administration. (D) Image analysis of Hamp2-labeled area at 1 and 26 weeks after tamoxifen administration. (E) Labeled clone size frequency in Hamp2-CreER livers at 1, 26, and 52 weeks after tamoxifen administration. (F) Individual clone size distribution in Hamp2-CreER livers at 1, 26, and 52 weeks after tamoxifen administration. The mean clone size (horizontal black line) increased from 1.38 ± 0.37 to 3.81 ± 0.35 to 4.74 ± 0.48 cells at 1, 26, and 52 weeks, respectively (mean ± SD). (G) Statistically representative cartoon of clone size changes over time. (H) Pie chart showing how different-sized clones contributed to the total number of labeled cells. ***P < 0.001. Error bars indicate SD for all panels except (B), where SEM is used.