Each diagonal panel represents individual distribution of each of four parameters among composition effect profiles promoting irreversible (green), reversible (orange), and no differentiation (black) strategies. Each non-diagonal panel represents a pairwise co-distribution of these parameters. Irreversible (green) strategies are promoted at small contribution thresholds and intermediary maximal benefit . Also, either the contribution synergy must be large, or the saturation threshold should be small – see main text for detailed discussion. Reversible (orange) strategies require large – there the benefits of having a large number of soma-role cells outweighs costs paid by frequent differentiation. Due to the fast accumulation of soma-role cells, reversible strategies tolerate larger than irreversible. Reversible exhibit the same restrictions with respect to as irreversible and are insensitive to . For this figure, 3000 composition effect profiles were investigated with costs and .