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Lineage P.4 defining mutations.
| Genomic region | Nucleotide | Amino acid |
|---|---|---|
| A136G | – | |
| ORF1a | G1811A | A516T |
| ORF1a | C3177T | P971L |
| ORF1a | C9693T | A3143V |
| ORF1a | T9867C | L3201P |
| ORF1a | C11450A | Q3729K |
| ORF1a | C12008T | L3915F |
| ORF1a | C12880T | – |
| ORF1b | A15932G | Y822C |
| ORF1b | G18756T | – |
| ORF1b | C20436T | – |
| S | G21987T | G142V |
| S | T22054G | N164K |
| S | C22079A | Q173K |
| S | T22917G | L452R |
| S | C23673T | S704L |
| S | A23720G | I720V |
| ORF3a | G25540A | V50I |
| ORF8 | C28253T | – |
| A28271C | – | |
| N | C28932T | A220V |
