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. 2021 Oct 6;12:729244. doi: 10.3389/fmicb.2021.729244

TABLE 1.

Wood and forest litter characteristics in unpolluted (UP), moderately polluted (MP), and heavily polluted (HP) areas.

Parameter Wood
Forest litter close to CWD
Forest litter far from CWD
UP area MP area HP area UP area MP area HP area UP area MP area HP area
Number of fir/spruce logs 19/11 12/18 18/12
Moss coverage, % of observed area 90.17 ± 12.21 66.83 ± 25.55 61.07 ± 29.97
Number of logs with herbs/tree seedlings 21/5 16/19 0/18
Circumference, cm 68.50 ± 22.34 68.83 ± 18.7 63.07 ± 28.51
Residence time, years 31.80 ± 12.47b 40.9 ± 13.14a 42.33 ± 11.54a
RP, J/cm3 3.77 ± 2.03a 2.52 ± 2.14b 3.13 ± 2.21ab
Submergence,% 25.14 ± 18.67b 38.19 ± 15.92a 41.13 ± 19.29a
Moss coverage, % 90.17 ± 12.21a 68.97 ± 22.72b 61.07 ± 29.97b
Moisture content 3.76 ± 1.14b 3.82 ± 1.09b 3.05 ± 0.86a 2.21 ± 0.51 2.14 ± 0.42 2.03 ± 0.49 1.95 ± 0.35 1.95 ± 0.40 1.86 ± 0.47
Ash content, % 2.23 ± 1.18 1.52 ± 0.80 1.66 ± 0.76 21.46 ± 7.47a 13.84 ± 3.45b 19.56 ± 5.58a 21.77 ± 8.33a 15.43 ± 5.43b 21.99 ± 8.69a
Hydrolytic acidity, mkM/100 g 52.05 ± 14.41b 59.63 ± 16.06ab 62.41 ± 11.38a 36.66 ± 7.72c 57.03 ± 6.70b 65.72 ± 6.24a 34.16 ± 9.31c 56.73 ± 6.49b 65.66 ± 7.04a
pH 3.62–5.73a 3.77–5.1ab 3.54–4.81b 4.94–6.35a 4.58–5.38b 4.32–5.33b 4.95–6.65a 4.56–5.45b 4.43–5.53c
P2O5, mg/100 g 16.78 ± 5.29 15.03 ± 7.65 20.71 ± 15.16 41.06 ± 9.80b 34.34 ± 5.19b 77.62 ± 23.67a 42.43 ± 8.78b 34.03 ± 3.72b 69.27 ± 22.15a
Total nitrogen, % 0.31 ± 0.14 0.22 ± 0.09 0.29 ± 0.17 1.63 ± 0.23 1.68 ± 0.27 1.61 ± 0.24 1.54 ± 0.25 1.69 ± 0.25 1.58 ± 0.27
Concentrations of acid-soluble metal forms, μg/g
Cu 6.32 ± 3.74c 46.02 ± 41.39b 188.13 ± 161.52a 23.50 ± 6.63c 929.80 ± 402.51b 2,406.89 ± 904.42a 22.80 ± 6.17c 959.42 ± 306.07b 2,405.10 ± 883.93a
Zn 48.24 ± 14.23c 125.90 ± 56.44b 240.68 ± 140.54a 232.94 ± 95.50b 574.85 ± 193.29a 554.55 ± 179.37a 193.00 ± 78.53b 560.38 ± 226.88a 537.28 ± 217.76a
Cd 0.53 ± 0.20c 2.06 ± 1.01b 4.27 ± 2.45a 2.12 ± 0.66b 10.73 ± 3.53a 10.63 ± 4.13a 1.92 ± 0.45b 10.19 ± 4.12a 10.04 ± 4.88a
Pb 2.81 ± 2.31c 20.55 ± 18.71b 67.69 ± 58.34a 47.92 ± 20.19c 1,013.41 ± 307.84b 1,959.55 ± 625.77a 42.19 ± 14.97c 1,031.18 ± 276.62b 1,945.26 ± 651.70a
Fe 51.00 ± 49.17b 58.22 ± 59.69b 191.82 ± 188.68a 985.39 ± 380.89b 1,708.40 ± 946.31b 5,856.60 ± 3,463.11a 1,232.81 ± 784.95b 1,894.29 ± 889.79b 5,919.21 ± 4,261.33a
Concentrations of exchangeable metal forms, μg/g
Cu 0.91 ± 0.71c 4.94 ± 6.01b 23.37 ± 24.95a 0.68 ± 0.19c 21.55 ± 11.70b 128.26 ± 74.66a 0.64 ± 0.18c 22.37 ± 10.52b 134.13 ± 96.83a
Zn 23.61 ± 8.84c 75.21 ± 34.03b 137.78 ± 88.23a 32.48 ± 11.98b 203.07 ± 48.23a 264.57 ± 78.57a 24.85 ± 10.14b 195.98 ± 58.37a 240.55 ± 72.23a
Cd 0.40 ± 0.20c 2.17 ± 1.13b 4.13 ± 2.77b 0.81 ± 0.26b 7.85 ± 2.06a 9.29 ± 3.59a 0.74 ± 0.25b 7.53 ± 2.25a 8.26 ± 3.04a
Pb 0.86 ± 0.31c 2.78 ± 2.09b 7.47 ± 8.10a 0.78 ± 0.29c 16.03 ± 9.35b 59.14 ± 26.02a 0.81 ± 0.31c 16.38 ± 8.71b 54.88 ± 30.72a
Fe 4.35 ± 2.84b 5.45 ± 5.21b 16.45 ± 21.70a 1.91 ± 0.93c 18.33 ± 20.88b 95.02 ± 91.89a 2.23 ± 2.55c 21.51 ± 26.39b 103.72 ± 95.63a

Mean values and standard deviations are shown; minimal and maximal values are provided for pH; n = 30 samples. Different letters depict statistically significantly different mean values between areas for a particular substrate (Tukey’s test, α = 0.05). CWD, coarse woody debris.