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Design of two-strain consortium that forms a synthetic extracellular protein matrix (sEPM). Schematic of C. crescentus secretor strain capable of exporting hydrogel proteins containing SpyCatcher covalent binding motif via a type I secretion system (A), C. crescentus displayer strain capable of binding SpyCatcher-hydrogel proteins to engineered RsaA-SpyTag S-layer at high density (B), and encapsulation of displayer cells in the secreted hydrogel protein forming a sEPM (C).
