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. Author manuscript; available in PMC: 2022 Dec 1.
Published in final edited form as: Am J Med Genet A. 2021 Aug 2;185(12):3784–3792. doi: 10.1002/ajmg.a.62447

Table 1.

Information on clinical phenotype and identified potentially pathogenic variants in novel VACTERL candidate genes.

Family Phenotypes Gene Pathway/
Complex
Mode of
transmission
Nucleotide
change
State Amino acid
change
Evolutionary
conservationA
PP2
SIFT
MT
CADD
SNP
ID
gnomad
B
ACMG
C
Segregation
381_501 A, C, R B9D1 Sonic hedgehog signaling, cilia organization recessive c.324G>A het p.(Val89Met) D. rerio 0.982 Del. D.C. 27.9 rs777500443 0/2/251 074 VUS (PM2, PP3) Variant inherited from healthy father
  B9D1 Sonic hedgehog signaling, cilia organization recessive c.71_72del het p.(Pro24fs) / / rs748661746 0/2/150 750 Likely pathogenic (PVS1 ,PM2) NA*
794_
501
V, A, R FREM1 Fraser complex recessive c.1408C>T het p.(Leu470Phe) M. musculus 0.359 Del. D.C. 23.1 rs1418448401 0/0/245 792 VUS (PM2, PP3) Variant inherited from healthy father
  FREM1 Fraser complex recessive c.56C>T het p.(Ala19Val) / 0.001 Tol. Poly. 7.947 rs368794455 0/6/232 056 VUS (PM2) Variant inherited from healthy mother
358_
501
V, R ZNF157 Kruppel family linking motifs, DNA transcription regulation NA c.764T>C hemi p.(Phe255Ser) X. tropicalis 0.999 Del. Poly. 10.51 / / VUS (PM2, PP3) Variant inherited from healthy mother
GE_
052
V, R ZNF157 Kruppel family linking motifs, DNA transcription regulation NA c.667T>G hemi p.(Cys223Gly) D. rerio 0.999 Del. D.C. 15.48 / / VUS (PM2, PP3) Variant inherited from healthy mother
395_
501
V, A, R, L SP8 Wnt, Shh and Bmp signaling NA c.1510C>T het p.(Arg504Cys) C. intestinalis 0.988 Del. D.C. 31 rs368503334 0/78/131 992 VUS (PM2, PP3) Variant inherited from healthy mother
SP8 Wnt, Shh and Bmp signaling NA c.417C>G het p.(Phe139Leu) D. rerio 0.086 Tol. D.C. 17.23 rs779330461 2/218/103 172 VUS (PM2, PP3) Variant inherited from healthy father
605_
501
V, A, C, R ACOT9 Fatty Acyl-CoA Biosynthesis recessive  c.953G>A hemi p.(Arg318Gln) D. rerio 0.812 Del. D.C. 32 / / VUS (PM2, PP3) Variant inherited from healthy mother
706_
501
A, R, L TTLL11 Spindle and cilia microtubules organization recessive c.1540-8C>A het p.? / / / / VUS (PM2) Variant inherited from healthy mother
    TTLL11 Spindle and cilia microtubules organization recessive c.1298C>G het  p.(Thr433Ser) D. melanogaster 0.997 Del. D.C. 25.9 rs187536422 0/2/251 496 VUS (PM2, PP3) Variant inherited from healthy father

Information on phenotype: A, anorectal malformation; C, cardiac malformation; L, limb anomalies; R, renal anomalies; V, vertebral malformation

CADD, Combined Annotated Dependent Depletion; D.C., disease-causing; Del., deleterious; del; deletion; fs, frameshift; gnomAD, Genome Aggregation Database; het, heterozygous; MT, Mutation Taster; NA, not available; PP2, PolyPhen 2; SIFT, Sorting intolerant from tolerant

A

Evolutionary conservation was assessed over 8 species: M. muscularis, Mus musculus; G. gallus, Gallus gallus; X. tropicalis, Xenopus tropicalis; D. rerio, Danio rerio; C. elegans, Caenorhabditis elegans; C. intestinalis, Ciona intestinalis; D. melanogaster, Drosphilia melanogaster; S. cerevisiae, Saccharomyces cerevisiae.

B

Variant frequencies listed for homozygous/ (if applicable) hemizygous/ heterozygous/ total alleles detected in the population.

C

ACMG American College of Medical Genetics guidelines classification as pathogenic, likely pathogenic, or variant of uncertain significance (VUS) (Richards et al., 2015).

*

no maternal DNA available; according to frequencies in gnomAD decided to be compound heterozygous.