TABLE 2.

Diversification rate models tested by GeoSSE/GeoHiSSE

	Model description	Range effect	Hidden states	AIC weights	AIC (%)
M1	GeoSSE, no range effect, no hidden state effect, null model	No	No	2.071081e−05	0.002
M2	GeoSSE, range effect, no hidden state effect (original GeoSSE model)	Yes	No	2.432576e−01	24.3
M3	GeoHiSSE, no range effect, hidden state effect	No	Yes	2.802921e−06	<0.001
M4	GeoHiSSE, both range and hidden state effect	Yes	Yes	7.567189e−01	75.6

Abbreviation: AIC, Akaike information criterion.