Spectrograms illustrating the hierarchical structure of humpback whale song. A single unit (trumpet) and a single phrase from Theme 25a are shown in the top panel. Theme 25a units from the single phrase in the top panel are as follows: short ascending moan, grunt, grunt, grunt, grunt, grunt, grunt, short ascending moan, trumpet, squeak, trumpet, squeak, trumpet. The repetition of phrases and the sequential singing of themes are shown in each of the subsequent panels. Spectrograms were 2048 point FFT, Hann window, 31 Hz resolution, and 75% overlap, generated in Raven Pro 1.4. Reprinted with permission from Garland et al. [48]. The devil is in the detail: quantifying vocal variation in a complex, multi-levelled, and rapidly evolving display. J. Acoust. Soc. Am.
142, 460–472 [48]. Copyright 2017, Acoustic Society of America.