1 |
Elytral disc broadly, deeply transversely impressed with a saddle-like depression from scutellum to declivital base; declivity deeply sulcate, its margins costate; elytral bases slightly emarginated to accommodate a prominent and dense mycangial tuft |
2
|
– |
Elytral disc either medially impressed and appearing humped, or flat, or broadly convex; declivity round or flat or convex; elytral bases not emarginated and mycangial tuft minute, either not readily apparent or lightly setose |
3
|
2 |
Elytral base emargination broadly recurved; elytral disc shallowly depressed; declivital summit armed with a pair of large tubercles at interstriae 1; punctures on declivity distinct and broad; larger species, 3.0–3.1 mm |
cruralis
|
– |
Elytral base emargination broadly V-shaped; elytral disc deeply depressed, more deeper posteriorly; interstriae 1 on declivital summit without a pair of large tubercles; punctures on declivital face small and indistinct; smaller species, 2.12 mm |
exsculptus
|
3 |
Posterolateral margin of elytra smoothly rounded, without any indication of a costa or carina |
4
|
– |
Posterolateral margin of elytra marked by an elevated costa or carina |
5
|
4 |
Elytral lateral margin tapering posteriorly from the middle to apex; elytral apex acute; minute species, 1.12–1.24 mm |
nanus
|
– |
Elytral lateral margin subparallel, broadly tapering posteriorly; apex broadly rounded, not acute; larger species, 1.60 mm |
globodeclivis sp. nov.
|
5 |
Declivity obliquely truncate; posterolateral declivital margin rounded and denticulate |
6
|
– |
Declivity rounded; posterolateral declivital margin costate or carinate, with or without granules |
9
|
6 |
Declivital interstriae 2 and 3 strongly laterally broadened from base to declivital midpoint and then narrowing towards apex |
latesalebrinus
|
– |
Declivital interstriae parallel from base to apex, never laterally broadened |
7
|
7 |
Denticles on declivital summit and margins larger and more sharply acute than those on declivital face |
kirishimanus
|
– |
Denticles on declivital summit of equal size and shape as those on declivital face |
8
|
8 |
Denticles on declivital summit as dense as those on declivital face; declivital face opalescent, subshiny |
nudibrevis
|
– |
Denticles on declivital summit denser than those on declivital face; declivital face shagreened, dull |
perparvus
|
9 |
Larger, 2.55–2.95 mm |
10
|
– |
Smaller, 1.2–2.1 mm |
12
|
10 |
Stout, 1.93–2.19× as long as wide; elytral posterolateral margin strongly carinate and unarmed |
fulvulus
|
– |
Elongate, 2.5–2.9× as long as wide; elytral posterolateral margin costate and granulate |
11
|
11 |
Declivital strial punctures very large, distinct |
chrysophylli
|
– |
Declivital strial punctures small, indistinct |
corporaali
|
12 |
Declivity with granules, denticles or tubercles distinctly less abundant than strial punctures |
13
|
– |
Declivity with abundant granules or denticles, at least as abundant as strial punctures |
19
|
13 |
Elytral disc shallowly transversely impressed with a saddle-like impression |
14
|
– |
Elytral disc without a depression |
15
|
14 |
Discal impression deeper, antero-posteriorly narrower, with steeper anterior and posterior slopes, strial punctures on impression with rounded granules; interstrial spines on disc behind impression stronger and backwardly hooked |
sagmatus
|
– |
Discal impression shallower, antero-posteriorly broader, with gentler anterior and posterior slopes, strial punctures on impression without granules; interstrial tubercles on disc behind impression moderate with rounded apices pointing dorsally |
undulatus
|
15 |
Declivital tubercles uniformly sized |
16
|
– |
Declivital tubercles not uniformly sized, 1–2 pairs of larger tubercles on declivital interstriae 3 |
17
|
16 |
Posterolateral declivital margin carinate, unarmed |
alpha
|
– |
Posterolateral declivital margin costate and denticulate |
bucolicus sp. nov.
|
17 |
Declivity steeply rounded, declivital interstriae 1 bearing 1–2 small tubercles on declivital face; larger, 1.9–2.0 mm |
recidens
|
– |
Declivity gradually rounded, declivital interstriae 1 unarmed on declivital face; smaller, 1.66–1.76 mm |
18
|
18 |
Declivital interstriae 3 with two pairs of prominent denticles; posterior half of elytral disc rugose with deeply impressed striae and costate interstriae, interstriae bearing strong incurved spines that become more prominent toward declivital summit |
serratus sp. nov.
|
– |
Declivital interstriae 3 with one pair of prominent tubercles; posterior half of elytral disc smooth, striae and interstriae flush, interstriae with a few small tubercles at summit |
bidentatus sp. nov.
|
19 |
Elytral disc convex on basal 1/3, appearing humped in lateral view |
20
|
– |
Elytral disc flat, never appearing humped |
22
|
20 |
Declivital interstriae densely covered with short semi-erect scales |
kadoyamaensis
|
– |
Declivital interstriae densely covered with long fine, erect hair-like setae |
21
|
21 |
Antennal club flat, type 3 with two sutures visible on apical 1/3 of posterior face; larger, declivity smooth, shiny; larger, 1.95–2.0 mm and more elongate, 2.79–2.86× as long as wide |
minax
|
– |
Antennal club obliquely truncate, type 2 with segment 1 almost covering posterior face (Fig. 2); one suture visible on posterior face near apex; declivity shagreened, dull; smaller, 1.8–1.9 mm and less elongate, 2.57–2.71× as long as wide |
nugax
|
22 |
Antennal club flat, type 3 with two sutures visible on apical 1/3 of posterior face (Fig. 3
|
diversicolor
|
– |
Antennal club obliquely truncate, type 2 with segment 1 almost covering posterior face; one suture visible on posterior face near apex |
23
|
23 |
Declivital interstrial granules dispersed, separated by the width of at least three granules; posterolateral margin of declivity weakly carinate and granulate; interstrial vestiture consisting of short semi-erect bristles, shorter in length than the width of an interstria; smaller, 1.2–1.7 mm |
pometianus
|
– |
Declivital interstrial granules dense, separated by the width of one granule; posterolateral margin of declivity strongly carinate; interstrial vestiture consisting of long semi-erect hair-like setae, longer in length than the width of an interstria (easily abraded); larger, 1.8–2.0 mm |
quercicola
|