Table 3.
Phenotypic characters useful for the taxonomy of Tityus. (***) Species here transferred to the indicated subgenus; ventral macrosetae distribution on telotarsi I–IV: Type I= tuft of irregularly distributed macrosetae. Type II= two discrete ventrosubmedian rows of macrosetae. Abbreviations: BML, basal middle lamellae; D, dilated; NA, not applicable; ND, not dilated; PBP, pectinal basal piece.
| Species | Subgenus | Species Group | Telotarsal setae | Females | |
|---|---|---|---|---|---|
| PBP gland | BML | ||||
| Isometrusmaculatus (DeGeer, 1778) | NA | NA | Type II | Absent | ND |
| Tityusargentinus Borelli, 1899 | Tityus | T.bolivianus | Type I | Absent | D= semicircular |
| Tityusbahiensis (Perty, 1833) | Tityus | T.bahiensis | Type II | First 2/3 of the anterior region | ND |
| Tityusblaseri Mello-Leitão, 1931 | Tityus | T.bahiensis | Type II | More than first 2/3 of the anterior region | ND |
| Tityusbrazilae Lourenço & Eickstedt, 1984*** | Tityus | T.bahiensis | Type II | First 2/3 of the anterior region | ND |
| Tityuscarrilloi Ojanguren-Affilastro, 2021 | Tityus | T.trivittatus | Type II | First 2/3 of the anterior region | ND |
| Tityuscarvalhoi Mello-Leitão, 1945 | Tityus | T.trivittatus | Type II | First 2/3 of the anterior region | ND |
| Tityuscharreyroni Mello-Leitão, 1933 | Tityus | T.bahiensis | Type II | First 2/3 of the anterior region | ND |
| Tityusclathratus C. L. Koch, 1844 | Archaeotityus | T.clathratus | Type I | First anteromedian third | ND |
| Tityusconfluens Borelli, 1899 | Tityus | T.trivittatus | Type II | First 2/3 of the anterior region | ND |
| Tityuscurupi Ojanguren-Affilastro et al. 2017 | Tityus | T.bolivianus | Type II | Absent | D= suboval |
| Tityusforcipula (Gervais, 1843) | Atreus | T.forcipula | Type II | Absent | D= suboval |
| Tityusobscurus Gervais, 1843 | Atreus | T.obscurus | Type I | Absent | D= semicircular |
| Tityuspanguana Kovařík et al. 2015 | Tityus | T.bolivianus | Type II | Absent | D= semicircular |
| Tityuspotameis Lourenço & Giupponi, 2004 | Tityus | T.trivittatus | Type II | First 2/3 of the anterior region | ND |
| Tityussastrei Lourenço & Flórez, 1990*** | Atreus | NA | Type II | Absent | D= semicircular |
| Tityusserrulatus Lutz & Melo, 1922 | Tityus | T.stigmurus | Type II | First anteromedian third | ND |
| Tityussoratensis Kraepelin, 1912 | Tityus | T.bolivianus | ? | Absent | D= semicircular |
| Tityusspelaeus sp. nov. | Tityus | T.trivittatus | Type II | More than first 2/3 of the anterior region | ND |
| Tityusstigmurus (Thorell, 1876) | Tityus | T.stigmurus | Type II | First anteromedian third | ND |
| Tityustrivittatus Kraepelin, 1898 | Tityus | T.trivittatus | Type II | First 2/3 of the anterior region | ND |
| Tityusuruguayensis Borelli, 1901 | Tityus | T.bolivianus | Type I | Absent | D= semicircular |