To self-organize multiple functionally diverse F-actin networks simultaneously from a common pool of components, both animal and fission yeast cells must (A) properly allocate actin monomers between competing networks, and (B) sort unique subsets of actin-binding proteins, which establish the structure and dynamics of each network. (C) ABPs compete and cooperate with each other, leading to a potential ABP hierarchy, where upstream proteins either recruit or inhibit the access of downstream proteins to specific F-actin networks.