TABLE 1.

Reference for metrics and assays.

Behavior metrics	Name	Measure
NTA	Net turn angle	Net sum of leftward and rightward angular movement
TTA	Total turn angle	Absolute sum of all angular movement
BR	Bias ratio	NTA/TTA ratio. Proportion of directional movement
MI	Match index	Proportion of trials matching direction of first trial
PI	Performance index	Average of binarized turn directions (0 = left; 1 = right)
Behavior assays	Name	Measure
4×	NA	Paired 30 s Light ON and OFF recordings, repeated 4×
8×	NA	Paired 30 s Light ON and OFF recordings, repeated 8×
q4×	Quad 4×	Four repeated ‘4×’ recordings separated by 10 min