Table 3.

Neuron, network, and simulation parameters.

Neuron and network parameters
Populations and external in-degree
Symbol		Value								Description
X		L2/3E	L2/3I	L4E	L4I	L5E	L5I	L6E	L6I	Population name
N X		20, 683	5, 834	21, 915	5, 479	4, 850	1, 065	14, 395	2, 948	Size
K X, ext		1, 600	1, 500	2, 100	1, 900	2, 000	1, 900	2, 900	2, 100	External in-degree
In-degrees in fixed in-degree models
K YX		from X
		L2/3E	L2/3I	L4E	L4I	L5E	L5I	L6E	L6I
	L2/3E	2,200	1,080	980	468	160	0	110	0
	L2/3I	2,991	861	704	290	381	0	61	0
	L4E	160	35	1,118	795	33	1	668	0
to Y	L4I	1,481	17	1,814	954	17	0	1,609	0
	L5E	2,189	375	1,136	32	421	497	297	0
	L5I	1,166	160	571	13	301	405	125	0
	L6E	326	39	468	92	286	22	582	753
	L6I	767	6	75	3	137	9	980	460
Total number of synapses in fixed total number models
S YX		from X
		L2/3E	L2/3I	L4E	L4I	L5E	L5I	L6E	L6I
	L2/3E	45,499,804	22,323,576	20,253,647	9,670,918	3,293,577	0	2,271,403	0
	L2/3I	17,443,694	5,018,762	4,105,338	1,690,073	2,221,212	0	353,460	0
	L4E	3,503,669	756,561	24,482,849	17,413,575	714,524	7,002	14,624,431	0
to Y	L4I	8,114,253	92,831	9,933,537	5,223,271	87,836	0	8,810,905	0
	L5E	10,613,575	1,817,058	5,507,804	151,900	2,040,738	2,407,889	1,438,969	0
	L5I	1,241,436	169,424	607,666	12,851	319,601	430,443	132,414	0
	L6E	4,681,225	556,108	6,727,569	1,320,233	4,112,224	305,028	837,2649	10,827,677
	L6I	2,260,836	17,207	220,032	8,078	401,637	25,217	2,888,426	1,354,319
Connection parameters and external input
Symbol			Value			Description
${\overline{w}}_{\infty }$			87.81 pA			Reference synaptic strength. All synapse weights are measured in units of ${\overline{w}}_{\infty }$.
g YX						Relative synaptic strengths:
			1			X ∈ {L2/3E, L4E, L5E, L6E}
			−4			X ∈ {L2/3I, L4I, L5I, L6I}, except for:
			2			(X, Y) = (L4E, L2/3E)
Δw∞, YX			$0.1·g_{YX}·{\overline{w}}_{\infty }$			Standard deviation of weight distribution
${\overline{d}}_{\mathrm{\text{E}}}$			1.5 ms			Mean excitatory delay
${\overline{d}}_I$			0.75 ms			Mean inhibitory delay
ΔdX			$0.5·{\overline{d}}_X$			Standard deviation of delay distribution
νext			8 s−1			Rate of external input with Poisson interspike interval statistics
w ext			${\overline{w}}_{\infty }$			Synaptic strength of external input
LIF neuron model
Symbol			Value			Description
C m			250 pF			Membrane capacitance
τm			10 ms			Membrane time constant
E L			−65 mV			Resistive leak reversal potential
V θ			−50 mV			Spike detection threshold
V reset			−65 mV			Spike reset potential
τref			2 ms			Absolute refractory period after spikes
τs			0.5 ms			Postsynaptic current time constant