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. 2022 Jan 25;12:1322. doi: 10.1038/s41598-022-05439-w

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© The Author(s) 2022

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/.

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Simplified principles of plasmodial Physarum polycephalum growth. (a) Growth of the plasmodium across seven-day observation period. (b) Illustrated depiction of plasmodial growth according to food source proximity. This behaviour leads to mesh clustering (phase one). (c) Illustrated depiction of mesh clustering, with mesh regions illustrated. (d) Illustrated depiction of network refinement and shortest-path optimization (phase two), where only a refined network remains and is illustrated. Illustrations overlaid atop true photographs.