Table 3.

Relationships in cooperative breeding Arctic fox families (Vulpes lagopus) that could be established through genetic analysis from tagged individuals

Year	Den ID	Group composition		Breeding females	Small rodent phase
		Females	Males
2002	Den 20	♀♀	♂♂	Mother and 1 yearling daughter	Decrease
2005	Den 20	♀♀♀	♂♂	Mother and 2 yearling daughters	Decrease
2007	Den 33	♀♀	♂	Mother and daughter (born 2005)	Increase
2011	Den 19	♀♀♀	♂♂	Mother and 2 yearling daughters	Peak
2014	Den 19	♀♀*	♂	Mother and daughter (born 2010)*	Increase
2015	Den 19	♀♀*	♂	Mother and daughter (born 2010)*	Peak
2015	Den 20	♀♀	♂	Mother and 1 yearling daughter	Peak
2017	Den 29	♀♀	♂	Mother and daughter (born 2015)	Increase
2017	Den 09	♀♀**	♂	Mother**	Increase
2017	Den 33	♀♀	♂	Mother and 1 yearling daughter	Increase

*One of the daughters born in 2010 stayed in the territory

**Not cooperative breeding, the only female visually observed at the den was the sister of the mother of the litter