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. 2022 Feb 21;13:834247. doi: 10.3389/fimmu.2022.834247

Table 2.

Properties of potential druggable targets in RA.

Leads from Multi-omics Pathology Cell Type Expression Signaling pathway Animal Model 3D Structures Clinical Status
MAP3K8/TPL2 (genomics, multiple evidences) Cell survival, production of inflammatory, degradative mediators, angiogenesis factors Synovial fibroblasts, myeloid cells, neutrophils MAPK pathways (ERK, p38 and JNK),
NFĸB signaling
Tpl2 knockdown in CIA mice;
inhibition in DSS-induced colitis mice model
5IU2
(catalytic site with adjacent selectivity pocket due to unique kinase domain structure)
SAR studies of preclinical molecules for RA; Clinical molecule for Ulcerative colitis
PADI4 (genomics, transcriptomics, epigenomics) NET formation, inflammation, regulation of T-cell mediated immune response Neutrophils, monocytes Citrullination pathway Inhibitor study in CIA murine model 4X8G, 4X8C (catalytic site) Preclinical molecules for RA, SLE
SHP2 (genomics and epigenomics) Survival and invasiveness of synovial lining FLS FLS, myeloid cells TNF-induced signaling through FAK and downstream MAPK (JNK, p38 and ERK) activation Heterozygous
PTPN11 deletion study in K/BxN serum transfer arthritis in mice
5EHR (allosteric site with hydrophobic subpocket can be targeted for selectivity) Preclinical non-competitive inhibitor for RA; Clinical molecules for solid tumors
PI3Kδ (epigenomics) Synovial hyperplasia, cell mobility and activation FLS in intimal lining, neutrophils, mast cells, B-cell, T-cell AKT and RAC signaling Inhibitor study in CIA rat model 4XE0 (Hydrophobic pocket adjacent to hinge region for selective lead design) SAR studies in RA; FDA approved inhibitor for cancer; Phase-2 inhibitors for asthma, COPD
HDAC3 (epigenomics) Inflammatory gene expression FLS, M1 macrophages, monocytes TLR signaling Only in vitro studies 4A69 (apo-structure, conformationally flexible site) SAR studies of preclinical molecules
BRD2/4 (epigenomics, genomics) Transcriptional activation of MMP, IL-6,8; angiogenesis FLS, macrophages, endothelial cells TNF-α/IL-1β/TLR signaling, VEGF-PAK1 signaling Inhibitor/siRNA study in CIA model 5EK9 (BRD2), 4MR4 (BRD4), BD2 domain selective; dual kinase/BRD inhibitors need to be explored SAR studies of preclinical molecules for RA; 11 clinical trial molecules in other diseases
CHKα (metabolic reprogram) Cell migration and proliferation, inflammation FLS, macrophages MAPK and PI3K/AKT signaling Inhibitor study in K/BxN serum transfer arthritis in mice 5AFV, 4DA5 (ATP binding site),
5W6O (allosteric site)
Preclinical molecules for RA; Clinical molecules for solid tumors
SPK1 (metabolic reprogram) Synovial hyperplasia, leucocyte infiltration inflammation FLS MAPK ERK, PI3K/AKT signaling Knockdown in CIA, AIA 3VZC, 4V24 (catalytic site with selective features of C4 domain) SAR studies of preclinical molecules for RA
HK2 (metabolic reprogram) Cell activation, invasive phenotype Synovial lining FLS Inducible glucose metabolism Deletion in K/BxN serum transfer arthritis model 5HG1, 5HFU (C-terminal catalytic pocket) SAR studies of preclinical molecules for cancer

MAPK, mitogen-activated protein kinase; SAR, structure-activity relationship; PADI4, peptidyl arginine deiminase 4; SLE, systemic lupus erythematosus; FLS, fibroblast-like synoviocytes; TNF, tumor necrosis factor; SHP2, Src homology-2 domain-containing protein tyrosine phosphatase-2; PI3Kδ, phosphatidylinositide 3-kinase delta; HDAC3, histone deacetylase 3; BRD2/4, bromodomain and extra-terminal proteins; CHKa, choline kinase alpha; SPK1, sphingosine kinase 1; HK2, hexokinase 2; CIA, collagen-induced arthritis; AIA, antibody induced arthritis.