Starvation conditions in yeast cause a major lifespan extension mediated in large part by the lack of amino acids and sugars. On one hand, amino-acid restriction causes the inactivation of TOR–Pkh–S6k signaling; on the other hand, low glucose levels promote reduced activity of the Ras–adenylate cyclase (Cyr1)–PKA pathway. Both the amino-acid and the sugar pathways converge on and inactivate the serine threonine kinase Rim15. This, in turn, contributes to the activation of stress-resistance transcription factors Gis1, which binds to post diauxic shift (PDS) motif, and Msn2 and Msn4, orthologs of mammalian early growth response protein 1 (EGR1)1,19, which bind to stress-responsive element (STRE) motif.