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. 2022 Mar 22;11:e72460. doi: 10.7554/eLife.72460

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© 2022, Mongiardino Koch et al

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Figure 1. — (A) Fellaster zelandiae, North Island, New Zealand (Clypeasteroida). (B) Large specimen: Peronella japonica, Ryukyu Islands, Japan; Small specimen: Echinocyamus crispus, Maricaban Island, Philippines (Laganina: Scutelloida). (C) Large specimen: Leodia sexiesperforata, Long Key, Florida; Small specimen: Sinaechinocyamus mai, Taiwan (Scutellina: Scutelloida). (D) Rhyncholampas pacificus, Isla Isabela, Galápagos Islands (Cassidulidae). (E) Conolampas sigsbei, Bimini, Bahamas (Echinolampadidae). (F)Apatopygus recens, Australia (Apatopygidae). (G) Hypotheses of relationships among neognathostomates. Top: Morphology supports a clade of Clypeasteroida + Scutelloida originating after the Cretaceous-Paleogene (K-Pg) boundary, subtended by a paraphyletic assemblage of extant (red) and extinct (green) ‘cassiduloids’ (Kroh and Smith, 2010). Bottom: A recent total-evidence study split cassiduloid diversity into a clade of extant lineages closely related to scutelloids, and an unrelated clade of extinct forms (Nucleolitoida; Mongiardino Koch and Thompson, 2021d). Divergence times are much older and conflict with fossil evidence. Cassidulids and apatopygids lacked molecular data in this analysis. Scale bars = 10 mm.