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. 2022 Mar 18;13:837756. doi: 10.3389/fphar.2022.837756

TABLE 2.

Significant compounds and pathways identified from the Reactome and Wilcoxon Rank sum test. Detailed information of the 83 compounds from the initial compound library that were shown to be significant in both the Reactome pathway analysis and wilcoxon rank sum test. The strictly standardized mean difference (SSMD) score measures the effect size and the variance amongst the triplicate larval samples for each compound. The brain health score (BHS) was defined as the logarithm of the covariance between the brain image and a template image. During the analysis pipeline, the SSMD and BHS scores were converted for directionality based on the pharmacological activity profile obtained from the Therapeutic Target database. The pathway names were outputted directly based on the target and activity profile from Reactome.

Compound Pathway name SSMD BHS Target Activity FDA status
Dexmedetomidine Adrenaline signalling through Alpha-2 adrenergic receptor 1.040 −2.928 ADRA2A AGONIST Approved
Guanabenz Acetate Adrenaline signalling through Alpha-2 adrenergic receptor 0.984 −0.868 ADRA2A AGONIST Approved
Noradrenaline Adrenaline signalling through Alpha-2 adrenergic receptor 0.855 −1.021 ADRA2A STIMULATOR Approved
Phentolamine Mesylate Adrenaline signalling through Alpha-2 adrenergic receptor −0.818 0.624 ADRA2A INHIBITOR Approved
Medetomidine Adrenaline signalling through Alpha-2 adrenergic receptor 0.777 −0.729 ADRA2A AGONIST Approved
Ivabradine HCl Adrenaline signalling through Alpha-2 adrenergic receptor 0.539 0.156 ADRA2A INHIBITOR Approved
Y-27632 2HCl Apoptosis 4.834 0.694 ROCK1 INHIBITOR
Oprozomib Apoptosis 1.558 0.221 PSMB8 INHIBITOR
Apoptosis Activator 2 Apoptosis 1.291 −3.112 CASP3 ACTIVATOR
Evodiamine Apoptosis −1.150 −0.525 BCL2 INDUCER
RKI-1447 Apoptosis 1.124 −0.097 ROCK1 INHIBITOR
Dynasore Apoptosis 0.913 −0.232 DNM1 INHIBITOR
PF-573228 Apoptosis 0.891 0.305 PTK2 INHIBITOR
Carfilzomib (PR-171) Apoptosis −0.801 −0.066 PSMD9 AGONIST Approved
ZSTK474 Cell surface interactions at the vascular wall 1.500 0.322 PIK3CA INHIBITOR
Dactolisib (BEZ235, NVP-BEZ235) Cell surface interactions at the vascular wall 0.904 0.571 PIK3CA INHIBITOR
RepSox Cell surface interactions at the vascular wall 0.746 −0.112 TGFB1 INHIBITOR
Dasatinib Cell surface interactions at the vascular wall 0.690 0.261 SRC INHIBITOR Approved
ML347 Cell surface interactions at the vascular wall 0.625 −0.090 TGFB1 INHIBITOR
CAL-101 (Idelalisib, GS-1101) Cell surface interactions at the vascular wall 0.590 0.360 PIK3CA INHIBITOR
Bosutinib (SKI-606) Cell surface interactions at the vascular wall 0.558 0.134 SRC INHIBITOR Approved
Ibuprofen (Dolgesic) COX reactions 1.124 0.217 COX INHIBITOR Approved
Mefenamic acid COX reactions 1.074 0.446 COX INHIBITOR Approved
Etodolac (Lodine) COX reactions 0.988 1.390 COX INHIBITOR Approved
Bromfenac COX reactions 0.778 1.053 COX INHIBITOR Approved
Nepafenac COX reactions 0.773 1.227 COX INHIBITOR Approved
Diclofenac Sodium COX reactions 0.694 0.428 PTSG2 INHIBITOR Approved
Ketorolac (ketorolac tromethamine) COX reactions 0.577 0.504 COX INHIBITOR Approved
Suprofen (Profenal) COX reactions 0.510 0.423 COX INHIBITOR Approved
Enzastaurin (LY317615) Depolymerisation of the Nuclear Lamina 0.522 0.610 PRKCB INHIBITOR
JTC-801 G-protein activation −1.223 −3.519 OPRM1 ANTAGONIST
Matrine ((+)-Matrine) G-protein activation 0.800 0.787 OPRM1 AGONIST
Naloxone HCl G-protein activation 0.564 0.964 OPRM1 AGONIST Approved
Tenovin-1 G2/M DNA damage checkpoint −1.612 −0.715 TP53 ACTIVATOR
VE-821 G2/M DNA damage checkpoint 0.923 0.332 ATM INHIBITOR
VE-822 G2/M DNA damage checkpoint 0.781 0.041 ATR ANTAGONIST
LY2608204 Glycolysis 1.145 1.058 GCK INHIBITOR
Clorsulon Glycolysis 0.907 0.518 GPM1 INHIBITOR
Vismodegib (GDC-0449) Hh mutants that don’t undergo autocatalytic processing are degraded by ERAD 1.472 0.739 SHH INHIBITOR Approved
PNU-120596 Highly calcium permeable nicotinic acetylcholine receptors −1.142 −3.868 CHRNA1 AGONIST
Tropicamide Highly calcium permeable nicotinic acetylcholine receptors 0.952 0.697 CHRNA1 INHIBITOR Approved
Darifenacin Highly calcium permeable nicotinic acetylcholine receptors 0.869 0.064 CHRNA1 INHIBITOR Approved
Pancuronium dibromide Highly calcium permeable nicotinic acetylcholine receptors 0.860 0.930 CHRNA1 INHIBITOR Approved
Gallamine triethiodide (Flaxedil) Highly calcium permeable nicotinic acetylcholine receptors 0.685 1.034 CHRNA1 INHIBITOR Approved
Adiphenine Highly calcium permeable nicotinic acetylcholine receptors 0.671 0.306 CHRNA1 INHIBITOR
Bethanechol chloride Highly calcium permeable nicotinic acetylcholine receptors 0.570 −0.201 CHRNA1 AGONIST Approved
Atropine sulfate monohydrate Highly calcium permeable nicotinic acetylcholine receptors 0.551 0.416 CHRNA1 INHIBITOR Approved
Cytisine Highly calcium permeable nicotinic acetylcholine receptors −0.520 −0.750 CHRNA4 AGONIST
Aliskiren hemifumarate Metabolism of Angiotensinogen to Angiotensins 1.540 1.1308 REN INHIBITOR Approved
Imidapril HCl Metabolism of Angiotensinogen to Angiotensins 0.938 0.5801 ACE INHIBITOR
Enalapril Maleate Metabolism of Angiotensinogen to Angiotensins 0.860 2.947 ACE INHIBITOR
Quinapril hydrochloride (accupril) Metabolism of Angiotensinogen to Angiotensins 0.707 0.385 ACE INHIBITOR Approved
Ramipril Metabolism of Angiotensinogen to Angiotensins 0.498 0.253 ACE INHIBITOR Approved
SB590885 Negative feedback regulation of MAPK pathway 1.699 0.738 RAF1 INHIBITOR
Selumetinib (AZD6244) Negative feedback regulation of MAPK pathway 1.098 0.172 MEK1 INHIBITOR
RAF265 (CHIR-265) Negative feedback regulation of MAPK pathway 0.886 0.537 RAF1 INHIBITOR
SL327 Negative feedback regulation of MAPK pathway 0.812 0.668 MEK1 INHIBITOR
Vemurafenib (PLX4032, RG7204) Negative feedback regulation of MAPK pathway 0.625 0.962 BRAF INHIBITOR Approved
Tanshinone IIA (Tanshinone B) Negative feedback regulation of MAPK pathway 0.547 1.823 MAP2K1 INHIBITOR
PD0325901 (PD325901) Negative feedback regulation of MAPK pathway 0.511 0.597 MEK1 INHIBITOR
IWR-1 (endo-IWR 1) PCP/CE pathway 2.195 0.994 WNT1 INHIBITOR
EHop-016 PCP/CE pathway 0.879 −0.273 RAC1 INHIBITOR
XAV-939 PCP/CE pathway 0.544 0.853 WNT1 INHIBITOR
Protionamide Peptide hormone metabolism 1.636 0.710 INHA INHIBITOR
Alogliptin Peptide hormone metabolism 0.988 0.720 DPP4 INHIBITOR Approved
TAK-875 Peptide hormone metabolism 0.733 1.320 gpr40 ANTAGONIST
SGC-CBP30 Regulation of Hypoxia-inducible Factor (HIF) by oxygen 2.312 1.079 DOT1L INHIBITOR
Rapamycin Regulation of TP53 Activity 1.679 1.365 MTOR INHIBITOR Approved
P22077 Regulation of TP53 Activity 1.145 0.694 USP7 INHIBITOR
ETP-46464 Regulation of TP53 Activity 1.085 0.023 MTOR INHIBITOR
Ridaforolimus Regulation of TP53 Activity 1.078 0.298 MTOR INHIBITOR
PP242 Regulation of TP53 Activity 0.896 0.892 MTOR INHIBITOR
KU-0063794 Regulation of TP53 Activity 0.618 1.254 MTOR INHIBITOR
PHT-427 Regulation of TP53 Activity 0.616 0.553 AKT1 INHIBITOR
Entinostat (MS-275) Regulation of TP53 Activity 0.574 0.524 HDAC1 INHIBITOR
AZD1152-HQPA (Barasertib) Regulation of TP53 Activity 0.517 1.01 AURKB INHIBITOR
Carprofen Respiratory electron transport 0.697 0.858 cox2 INHIBITOR Approved
Cilengitide Smooth Muscle Contraction 0.718 −0.104 ITGA1 INHIBITOR
(-)-Huperzine A Synthesis of PC 1.320 0.550 ACHE INHIBITOR
Odanacatib (MK 0822) Toll-Like Receptors Cascades −1.054 −0.098 CTSK AGONIST
EUK 134 Toll-Like Receptors Cascades 0.529 0.279 APP INHIBITOR
NMDA TP53 Regulates Metabolic Genes 1.691 0.619 NMDA AGONIST
BAM 7 TP53 Regulates Transcription of Genes Involved in G2 Cell Cycle Arrest 0.763 0.027 BAX INDUCER