Table 1.

Simulation parameters

Parameter	Definition	Value	Unit	Notes
a	fiber radius	4	nm	(38)
L	fiber length	$0.5,1$	μm	(39,40)
F	number of fibers	200–1,600
$L_d$	simulated volume’s extent	2–4	μm	cubic unit cell
μ	cytoplasm viscosity	0.1	Pa⋅s	100 $\times$ water (cytoplasm) (41)
κ	fiber bending stiffness	0.07	pN⋅$\mu$m2	17 μm persistence length (22)
$K_c$	CL spring stiffness	10	pN/μm	(42)
$\ell$	CL rest length	50	nm	(29)
$k_{\text{on}}$	CL first end binding rate	5	1/(μm⋅s)	(3)
$k_{\text{on},\text{s}}$	CL second end binding rate	50	1/(μm⋅s)	$k_{\text{on},\text{s}}\gg k_{\text{on}}$, not measured
$k_{\text{off}}$	CL (one end bound) unbinding rate	1	1/s	(43,44)
$k_{\text{off},\text{s}}$	CL (both ends bound) unbinding rate	1	1/s	$k_{\text{off},\text{s}}=k_{\text{off}}$
$c_w$	actin-binding site width	20	nm	(29)
$k_{B}T$	thermal energy	4 × 10−3	pN⋅$\mu$m	25°C
N	number of Chebyshev points	16		((21), Sec. 6.3.1)
$\text{Δ}{s}_u$	binding site spacing	0.026	μm	(3)
$\text{Δ}t$	time step size	${10}^{-4}$	s	limited by $K_c$