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. 2021 Aug 9;11(44):27226–27245. doi: 10.1039/d0ra07488h

Differential metabolites and their classes related to salt-alkali stress of C. deserticolaa.

No. Compounds Saline-alkali land vs. grassland Sandy land vs. saline-alkali land Class
Inflorescence (A1 vs. B1) Succulent stem (A2 vs. B2) Inflorescence (C1 vs. A1) Succulent stem (C2 vs. A2)
VIP Fold change Type VIP Fold change Type VIP Fold change Type VIP Fold change Type
CdM781 1-Methylhistidine 2.01 × 100 1.86 × 10−4 Down Amino acid derivatives
CdM182 l-Methionine methyl ester 2.11 × 100 7.96 × 10−5 Down 2.44 × 100 2.12 × 104 Up
CdM1039 2,3-Dimethylsuccinic acid 2.40 × 100 5.03 × 10−6 Down
CdM1064 Pyrrole-2-carboxylic acid 2.23 × 100 1.93 × 10−4 Down 2.13 × 100 5.18 × 103 Up
CdM781 1-Methylhistidine 2.39 × 100 1.95 × 104 Up
CdM192 3-Chloro-l-tyrosine 2.40 × 100 7.71 × 10−5 Down 2.67 × 100 5.12 × 10−5 Down 2.29 × 100 1.30 × 104 Up
CdM371 N-Phenylacetylglycine 2.50 × 100 4.44 × 104 Up 2.41 × 100 2.64 × 10−5 Down
CdM19 3-N-Methyl-l-histidine 2.65 × 100 1.72 × 105 Up
CdM393 Phe–Phe 2.51 × 100 1.61 × 10−4 Down 2.13 × 100 2.84 × 10−4 Down
CdM11 Histamine 2.28 × 100 1.92 × 104 Up
CdM799 l-Citrulline 2.15 × 100 5.08 × 10−5 Down Amino acids
CdM30 l-(+)-Arginine 2.25 × 100 1.26 × 10−5 Down
CdM837 l-Cysteine 2.58 × 100 3.18 × 104 Up 2.32 × 100 4.51 × 10−5 Down
CdM1087 Cyanidin O-diacetyl-hexoside-O-glyceric acid 1.84 × 100 1.54 × 103 Up 2.36 × 100 9.79 × 10−5 Down Anthocyanins
CdM317 Peonidin O-hexoside 2.17 × 100 2.11 × 104 Up
CdM230 Delphinidin 3-O-glucoside (mirtillin) 2.21 × 100 3.32 × 10−5 Down 2.45 × 100 1.94 × 104 Up
CdM357 Rosinidin O-hexoside 2.51 × 100 5.86 × 105 Up 2.76 × 100 2.90 × 10−6 Down
CdM267 Cyanidin 3-O-rutinoside (keracyanin) 2.92 × 100 7.03 × 10−8 Down 2.80 × 100 1.42 × 107 Up
CdM270 Petunidin 3-O-glucoside 2.72 × 100 2.63 × 104 Up
CdM255 Syringic acid O-feruloyl-O-hexoside 2.25 × 100 4.43 × 104 Up 2.38 × 100 2.64 × 10−5 Down Benzoic acid derivatives
CdM1059 2,4-Dihydroxybenzoic acid 2.47 × 100 2.20 × 10−6 Down
CdM733 4-Hydroxy-3,5-diisopropylbenzaldehyde 1.04 × 100 2.11 × 10−2 Down 2.41 × 100 1.64 × 105 Up
CdM989 Gallic acid O-hexoside 2.14 × 100 5.14 × 10−4 Down 2.05 × 100 1.94 × 103 Up
CdM630 3,4-Dihydrocoumarin 2.00 × 100 2.08 × 10−4 Down 2.30 × 100 1.58 × 10−4 Down 2.20 × 100 6.32 × 103 Up Coumarins
CdM528 6-Hydroxy-4-methylcoumarin 2.11 × 100 1.29 × 104 Up 2.42 × 100 2.28 × 104 Up 2.43 × 100 2.14 × 10−5 Down
CdM241 O-Feruloyl coumarin 2.11 × 100 1.21 × 104 Up 3.05 × 100 8.07 × 105 Up 2.27 × 100 7.67 × 10−5 Down
CdM359 Daphnetin 2.24 × 100 4.28 × 104 Up 1.06 × 100 1.43 × 101 Up 2.84 × 100 1.08 × 10−5 Down 1.07 × 100 1.16 × 10−1 Down
CdM426 N-Sinapoyl hydroxycoumarin 2.36 × 100 1.27 × 105 Up 2.57 × 100 1.35 × 10−5 Down 2.46 × 100 7.40 × 104 Up
CdM1086 Esculetin (6,7-dihydroxycoumarin) 1.25 × 100 9.05 × 10−2 Down 1.03 × 100 6.69 × 100 Up
CdM495 Hesperetin 7-O-neohesperidoside (neohesperidin) 2.07 × 100 8.68 × 10−5 Down Flavanone
CdM603 Eriodictyol 2.16 × 100 4.80 × 10−5 Down 2.28 × 100 2.46 × 104 Up
CdM464 Naringenin 7-O-neohesperidoside (naringin) 2.41 × 100 4.92 × 10−6 Down 2.54 × 100 2.51 × 105 Up
CdM1148 Hesperetin 5-O-glucoside 2.48 × 100 1.22 × 10−6 Down 3.08 × 100 8.20 × 105 Up
CdM731 Xanthohumol 2.06 × 100 7.96 × 10−4 Down
CdM1203 Naringenin 7-O-glucoside (prunin) 2.42 × 100 4.95 × 10−5 Down 2.31 × 100 2.02 × 104 Up
CdM1024 Chrysoeriol O-hexosyl-O-malonylhexoside 1.74 × 100 6.41 × 102 Up 2.19 × 100 3.04 × 10−4 Down Flavone
CdM1202 Tricin 7-O-hexoside 1.90 × 100 3.45 × 10−4 Down
CdM553 Tricin O-malonylhexoside 1.94 × 100 3.35 × 10−4 Down 2.40 × 100 4.16 × 10−5 Down
CdM506 Chrysoeriol O-hexosyl-O-hexoside 1.94 × 100 2.97 × 10−4 Down 2.06 × 100 1.35 × 103 Up
CdM1193 Tricin O-glucuronic acid 2.01 × 100 5.27 × 103 Up 2.21 × 100 1.44 × 10−4 Down
CdM1326 Acacetin 2.06 × 100 6.54 × 10−5 Down
CdM503 Syringetin 7-O-hexoside 2.10 × 100 6.86 × 10−5 Down 2.74 × 100 4.15 × 10−6 Down 2.62 × 100 2.41 × 105 Up
CdM381 Limocitrin O-hexoside 2.12 × 100 1.47 × 104 Up 2.44 × 100 1.89 × 104 Up
CdM353 Chrysoeriol O-glucuronic acid-O-hexoside 2.13 × 100 6.53 × 10−5 Down 2.31 × 100 1.52 × 10−4 Down 2.21 × 100 6.57 × 103 Up
CdM434 Acacetin O-glucuronic acid 2.25 × 100 1.50 × 10−5 Down 2.40 × 100 1.15 × 105 Up
CdM510 Apigenin 7-O-glucoside (cosmosiin) 2.37 × 100 4.97 × 10−6 Down 2.39 × 100 4.41 × 104 Up
CdM1253 Luteolin 2.38 × 100 2.54 × 10−6 Down 1.49 × 100 4.59 × 101 Up
CdM529 Selgin O-malonylhexoside 2.63 × 100 4.57 × 10−7 Down 2.47 × 100 3.67 × 10−5 Down 1.25 × 100 4.79 × 100 Up 2.36 × 100 2.72 × 104 Up
CdM492 Chrysoeriol 7-O-rutinoside 2.29 × 100 1.62 × 10−4 Down 2.19 × 100 6.18 × 103 Up
CdM515 Chrysoeriol O-glucuronic acid 2.34 × 100 3.67 × 10−5 Down 2.24 × 100 2.72 × 104 Up
CdM595 Chrysin O-hexoside 2.35 × 100 1.03 × 10−4 Down 2.25 × 100 9.68 × 103 Up
CdM594 Chrysin 5-O-glucoside (toringin) 2.36 × 100 8.14 × 10−5 Down 2.26 × 100 1.23 × 104 Up
CdM543 Apigenin O-malonylhexoside 2.37 × 100 9.35 × 10−5 Down 2.27 × 100 1.07 × 104 Up
CdM1186 Chrysoeriol O-rhamnosyl-O-glucuronic acid 2.45 × 100 1.11 × 10−5 Down 2.35 × 100 9.03 × 104 Up
CdM396 Selgin 5-O-hexoside 2.47 × 100 3.13 × 10−5 Down
CdM485 Syringetin 5-O-hexoside 2.74 × 100 3.93 × 10−6 Down 2.85 × 100 9.84 × 10−6 Down 2.62 × 100 2.54 × 105 Up
CdM1202 Tricin 7-O-hexoside 2.36 × 100 2.90 × 103 Up
CdM553 Tricin O-malonylhexoside 2.41 × 100 2.99 × 103 Up 2.29 × 100 2.41 × 104 Up
CdM506 Chrysoeriol O-hexosyl-O-hexoside 2.41 × 100 3.36 × 103 Up
CdM522 Spinacetin 2.48 × 100 1.20 × 10−4 Down
CdM1030 Luteolin O-hexosyl-O-hexosyl-O-hexoside 2.48 × 100 5.75 × 103 Up
CdM556 Chrysoeriol O-malonylhexoside 2.62 × 100 7.15 × 10−5 Down 2.30 × 100 5.96 × 10−5 Down
CdM1135 Tricin O-saccharic acid 1.03 × 100 1.43 × 10−1 Down
CdM1211 Baicalein-7-O-glucuronide (baicalin) 1.36 × 100 3.56 × 102 Up
CdM379 Tricin 7-O-hexosyl-O-hexoside 2.26 × 100 1.05 × 104 Up
CdM555 Tricetin O-malonylhexoside 2.40 × 100 4.16 × 10−5 Down
CdM632 Apigenin 2.36 × 100 4.82 × 104 Up
CdM471 Apigenin 7-rutinoside (isorhoifolin) 2.36 × 100 1.42 × 105 Up
CdM342 Tricin 5-O-hexosyl-O-hexoside 2.37 × 100 2.59 × 104 Up
CdM490 Apigenin 7-O-neohesperidoside (rhoifolin) 2.52 × 100 1.84 × 105 Up
CdM525 Apigenin 8-C-pentoside 2.06 × 100 8.40 × 103 Up 2.62 × 100 4.52 × 10−5 Down Flavone C-glycosides
CdM370 8-C-Hexosyl-luteolin O-pentoside 2.15 × 100 1.70 × 104 Up 2.69 × 100 3.80 × 10−5 Down
CdM1096 Eriodictyol C-hexoside 2.22 × 100 1.04 × 10−5 Down 2.45 × 100 2.36 × 10−5 Down 2.35 × 100 4.25 × 104 Up
CdM345 C-Hexosyl-apigenin O-feruloylhexoside-O-hexoside 2.26 × 100 1.95 × 10−5 Down 2.81 × 100 5.12 × 104 Up
CdM286 8-C-Hexosyl-hesperetin O-hexoside 2.32 × 100 6.94 × 10−6 Down 2.88 × 100 1.44 × 105 Up
CdM498 C-Pentosyl-apigenin O-p-coumaroylhexoside 2.35 × 100 1.26 × 105 Up 2.54 × 100 1.97 × 10−5 Down 2.90 × 100 9.61 × 10−6 Down 1.10 × 100 1.17 × 101 Up
CdM288 Chrysoeriol 6-C-hexoside 8-C-hexoside-O-hexoside 2.37 × 100 1.47 × 105 Up 2.32 × 100 5.87 × 10−5 Down
CdM424 8-C-Hexosyl-luteolin O-hexoside 2.67 × 100 3.01 × 10−7 Down
CdM366 6-C-Hexosyl-hesperetin O-hexoside 2.51 × 100 3.83 × 104 Up
CdM373 C-Hexosyl-chrysoeriol O-rutinoside 2.61 × 100 1.31 × 104 Up
CdM386 C-Hexosyl-chrysoeriol O-hexoside 2.72 × 100 3.93 × 10−5 Down 2.36 × 100 3.28 × 104 Up
CdM504 Isorhamnetin O-hexoside 1.19 × 100 4.70 × 10−1 Down 1.24 × 100 4.32 × 100 Up Flavonol
CdM1297 Isorhamnetin 1.89 × 100 5.43 × 10−4 Down 2.34 × 100 1.84 × 103 Up
CdM670 3,7-Di-O-methylquercetin 1.97 × 100 2.73 × 10−4 Down
CdM398 Isorhamnetin 3-O-neohesperidoside 2.01 × 100 1.86 × 10−4 Down 2.09 × 100 2.69 × 103 Up
CdM474 Kaempferide 2.01 × 100 6.30 × 103 Up
CdM1189 Dihydroquercetin (taxifolin) 2.02 × 100 8.04 × 10−5 Down
CdM1230 Aromadedrin (dihydrokaempferol) 2.06 × 100 9.40 × 10−5 Down
CdM1143 Kaempferol 3-O-rutinoside (nicotiflorin) 2.08 × 100 9.25 × 10−5 Down 2.58 × 100 1.08 × 104 Up
CdM1123 Kaempferol 3-O-robinobioside (biorobin) 2.10 × 100 8.40 × 10−5 Down
CdM308 Quercetin 5-O-malonylhexosyl-hexoside 2.22 × 100 1.95 × 10−5 Down 2.76 × 100 5.12 × 104 Up
CdM1164 Quercetin O-acetylhexoside 2.34 × 100 4.23 × 10−6 Down 2.91 × 100 2.37 × 105 Up
CdM1149 Quercetin 3-O-glucoside (isotrifoliin) 2.47 × 100 1.24 × 10−6 Down 3.07 × 100 8.08 × 105 Up
CdM1206 Isorhamnetin O-acetyl-hexoside 2.51 × 100 1.45 × 10−6 Down 2.12 × 100 5.77 × 10−4 Down 3.12 × 100 6.92 × 105 Up 2.03 × 100 1.73 × 103 Up
CdM457 Methyl quercetin O-hexoside 2.54 × 100 1.19 × 10−6 Down 3.15 × 100 8.42 × 105 Up
CdM1298 Di-O-methylquercetin 1.17 × 100 7.35 × 10−2 Down 1.08 × 100 1.16 × 101 Up
CdM466 Kaempferol 3-O-galactoside (trifolin) 2.42 × 100 1.78 × 104 Up 2.50 × 100 1.23 × 10−5 Down
CdM460 Quercetin 4′-O-glucoside (spiraeoside) 2.42 × 100 5.47 × 10−5 Down 3.18 × 100 2.24 × 106 Up 2.32 × 100 1.83 × 104 Up
CdM584 Icariin (kaempferol 3,7-O-diglucoside 8-prenyl derivative) 2.91 × 100 7.83 × 10−7 Down 1.39 × 100 3.60 × 101 Up
CdM497 Kaempferol 3-O-glucoside (astragalin) 2.51 × 100 9.11 × 103 Up
CdM1109 Dihydromyricetin 2.66 × 100 3.65 × 10−5 Down
CdM482 Isorhamnetin 5-O-hexoside 3.34 × 100 4.67 × 106 Up
CdM341 6-Hydroxymethylherniarin 1.95 × 100 3.40 × 103 Up 2.35 × 100 9.38 × 10−5 Down 2.58 × 100 9.90 × 10−5 Down Hydroxycinnamoyl derivatives
CdM433 Sinapic acid 2.06 × 100 1.26 × 10−4 Down
CdM508 p-Coumaraldehyde 2.36 × 100 1.22 × 105 Up 2.63 × 100 1.36 × 105 Up
CdM403 3-(4-Hydroxyphenyl)propionic acid 2.49 × 100 4.63 × 105 Up 2.73 × 100 2.90 × 105 Up 2.67 × 100 2.69 × 10−6 Down
CdM1054 Syringin 2.49 × 100 5.06 × 105 Up 1.05 × 100 1.31 × 10−1 Down
CdM1248 3,4,5-Trimethoxycinnamic acid 1.01 × 100 2.08 × 10−2 Down
CdM1271 4-Methoxycinnamic acid 1.09 × 100 8.21 × 10−2 Down
CdM1273 3,4-Dimethoxycinnamic acid 1.26 × 100 3.43 × 10−2 Down
CdM295 Gallic acid O-feruloyl-O-hexosyl-O-hexoside 2.43 × 100 5.88 × 10−5 Down 2.64 × 100 6.91 × 10−5 Down
CdM1080 1-O-Beta-d-glucopyranosyl sinapate 2.44 × 100 4.25 × 10−5 Down 2.34 × 100 2.35 × 104 Up
CdM508 p-Coumaraldehyde 2.61 × 100 4.17 × 10−6 Down
CdM403 3-(4-Hydroxyphenyl)propionic acid
CdM350 Homovanillic acid 2.83 × 100 1.70 × 10−6 Down
CdM330 Caffeic acid 3.20 × 100 3.61 × 106 Up 2.53 × 100 6.55 × 10−6 Down
CdM1158 Coniferyl alcohol 2.19 × 100 5.48 × 103 Up
CdM1209 6-Hydroxydaidzein 2.04 × 100 9.00 × 10−5 Down Isoflavone
CdM1325 Prunetin 2.06 × 100 6.26 × 10−5 Down
CdM583 Orobol (5,7,3′,4′-tetrahydroxyisoflavone) 2.16 × 100 2.56 × 10−5 Down 2.68 × 100 3.91 × 104 Up
CdM1213 2′-Hydroxydaidzein 2.19 × 100 3.31 × 10−5 Down
CdM623 Genistein (4′,5,7-trihydroxyisoflavone) 2.23 × 100 1.99 × 10−5 Down 2.77 × 100 5.02 × 104 Up
CdM465 Genistein 7-O-glucoside (genistin) 2.38 × 100 4.59 × 10−6 Down 2.42 × 100 5.27 × 104 Up
CdM1226 Formononetin 7-O-glucoside (ononin) 2.41 × 100 3.23 × 10−4 Down
CdM579 2′-Hydroxygenistein 2.92 × 100 2.08 × 105 Up
CdM1115 Glycitin 2.27 × 100 8.80 × 10−5 Down
CdM678 MAG (18 : 4) isomer 1 2.21 × 100 1.94 × 10−5 Down 2.52 × 100 1.78 × 10−5 Down 2.41 × 100 5.62 × 104 Up Lipids glycerolipids
CdM712 MAG (18 : 3) isomer 5 1.03 × 100 6.84 × 10−2 Down
CdM769 MAG (18 : 3) isomer 1 1.05 × 100 1.17 × 10−1 Down
CdM713 DGMG (18 : 1) 1.06 × 100 7.14 × 10−2 Down
CdM711 DGMG (18 : 2) isomer 2 1.07 × 100 7.23 × 10−2 Down
CdM767 MAG (18 : 3) isomer 4 1.08 × 100 1.05 × 10−1 Down
CdM168 Thymine 2.07 × 100 1.96 × 104 Up 2.75 × 100 1.97 × 10−5 Down Nucleotide and its derivatives
CdM161 8-Hydroxyguanosine 2.11 × 100 1.25 × 104 Up 2.59 × 100 9.92 × 10−5 Down
CdM183 5-Methyluridine 2.15 × 100 5.82 × 10−5 Down 2.36 × 100 8.80 × 10−5 Down
CdM179 Guanosine 3′,5′-cyclic monophosphate 2.15 × 100 5.33 × 10−5 Down 2.67 × 100 1.87 × 104 Up
CdM127 Uracil 1.12 × 100 1.51 × 10−1 Down
CdM61 Cytidine 5′-monophosphate (cytidylic acid) 2.09 × 100 3.28 × 10−4 Down 2.01 × 100 3.05 × 103 Up
CdM73 Inosine 2.38 × 100 2.40 × 103 Up
CdM18 2′-Deoxyinosine-5′-monophosphate 1.02 × 100 6.10 × 101 Up
CdM74 iP7G 1.23 × 100 1.03 × 10−1 Down
CdM585 Sebacate 1.98 × 100 2.58 × 10−4 Down 2.56 × 100 2.05 × 10−5 Down 1.08 × 100 1.34 × 10−1 Down Organic acids
CdM242 Sinapoyl malate 2.04 × 100 6.47 × 103 Up
CdM1091 p-Hydroxyphenyl acetic acid 2.25 × 100 7.00 × 10−6 Down
CdM1067 Mandelic acid 2.25 × 100 6.61 × 10−6 Down
CdM941 3-Hydroxybutyrate 2.26 × 100 1.73 × 10−5 Down
CdM328 4-Hydroxybenzoic acid 2.35 × 100 7.99 × 10−6 Down 2.44 × 100 9.33 × 104 Up
CdM1041 2-Methylglutaric acid 2.42 × 100 4.39 × 10−6 Down
CdM1040 Adipic acid 2.42 × 100 4.10 × 10−6 Down
CdM1042 5-Hydroxyhexanoic acid 1.12 × 100 1.26 × 10−1 Down 1.20 × 100 1.28 × 101 Up
CdM1051 2-Isopropylmalate 1.17 × 100 1.07 × 10−1 Down 1.25 × 100 1.57 × 101 Up
CdM1045 3-Hydroxyanthranilic acid 2.44 × 100 5.40 × 10−5 Down
CdM172 4-Acetamidobutyric acid 2.60 × 100 4.66 × 10−6 Down 2.49 × 100 2.15 × 105 Up
CdM577 N′,N′′-p-Coumaroyl-feruloyl putrescine 1.89 × 100 3.68 × 10−4 Down 2.34 × 100 2.72 × 103 Up Phenolamides
CdM394 N′,N′′-Diferuloylspermidine 2.11 × 100 8.25 × 10−5 Down 2.61 × 100 1.21 × 104 Up
CdM216 N-p-Coumaroyl putrescine 2.44 × 100 3.35 × 10−6 Down 3.03 × 100 2.99 × 105 Up
CdM217 N′-p-Coumaroyl putrescine 2.45 × 100 2.95 × 10−6 Down 3.04 × 100 3.39 × 105 Up
CdM365 N′,N′′-Di-p-coumaroylspermidine 2.58 × 100 8.26 × 10−7 Down 1.45 × 100 1.67 × 101 Up
CdM351 N′,N′′-feruloyl-caffeoylspermidine 2.25 × 100 2.17 × 10−4 Down
CdM1312 Gibberellin A4 (GA4) 1.72 × 100 1.57 × 10−3 Down Phytohormones
CdM1334 Gibberellin A15 1.86 × 100 1.77 × 103 Up 2.24 × 100 1.90 × 10−4 Down 2.42 × 100 2.70 × 10−4 Down 2.14 × 100 5.25 × 103 Up
CdM707 Gibberellin A9 2.09 × 100 5.87 × 10−5 Down 2.40 × 100 3.50 × 104 Up
CdM1292 (+)-Jasmonic acid (JA) 2.09 × 100 9.51 × 10−5 Down 2.60 × 100 1.05 × 104 Up
CdM606 Gibberellin A20 2.16 × 100 3.01 × 10−5 Down 2.67 × 100 3.32 × 104 Up 2.46 × 100 6.12 × 104 Up
CdM278 Indole carboxylic acid 2.16 × 100 2.03 × 104 Up 2.79 × 100 1.80 × 10−5 Down
CdM248 trans-Zeatin 9-O-glucoside 1.16 × 100 1.04 × 10−1 Down
CdM1001 5-O-Feruloyl quinic acid glucoside 1.78 × 100 8.36 × 102 Up Quinate and its derivatives
CdM1003 3-O-Feruloyl quinic acid glucoside 1.83 × 100 1.22 × 103 Up
CdM315 3-O-Feruloyl quinic acid 1.95 × 100 2.98 × 103 Up 2.22 × 100 3.51 × 103 Up 2.49 × 100 1.96 × 10−4 Down
CdM1108 3-O-p-Coumaroyl quinic acid 2.01 × 100 1.47 × 10−4 Down 2.25 × 100 1.90 × 10−4 Down 2.15 × 100 5.27 × 103 Up
CdM289 Chlorogenic acid (3-O-caffeoylquinic acid) 2.05 × 100 1.44 × 10−4 Down 2.54 × 100 6.92 × 103 Up
CdM1028 5-O-p-Coumaroyl shikimic acid O-hexoside 2.05 × 100 6.71 × 10−5 Down
CdM1079 3-O-p-Coumaroyl shikimic acid O-hexoside 2.07 × 100 9.14 × 103 Up 2.21 × 100 1.06 × 10−4 Down
CdM411 Chlorogenic acid methyl ester 2.49 × 100 5.36 × 105 Up
CdM1056 O-p-Coumaroyl quinacyl quinic acid O-hexoside 1.19 × 100 4.97 × 10−2 Down
a

Note: A1: inflorescence in saline-alkali land, A2: succulent stem in saline-alkali land, B1: inflorescence in grassland, B2: succulent stem in grassland, C1: inflorescence in sandy land, and C2: succulent stem in sandy land.