Partitioning of variation in the bacterial community composition of coral tissue (a) and skeleton (d) explained by host skeletal architecture, phylogeny, and reproductive mode. Adjusted R2 values are given. Canonical correspondence analysis (CCA) biplots represent the tissue (b) and skeletal (c) microbiome structures according to each explanatory variable. Arrows represent the quantitative explanatory variables skeletal architecture and phylogeny (PV1, PV2, and PV3), with arrowheads indicating the direction of increase. The categorical explanatory variables broadcast spawners, brooders, and mixed mode are positioned on the biplot according to their coordinates. All of the explanatory variables in both CCA biplots were highly significant (P < 0.001). In variation partitioning analysis (a, d), the total explained and unexplained variance can exceed 100%.