Phylogenetic tree of BAHD superfamily proteins. Sequence names are prefixed by a two-letter species code and colored by species: L. sativus (Ls), blue; A. thaliana (At), green; M. truncutula (Mt), purple; P. trichocarpa (Pt), brown; other species are colored black. The outer ring and clades are colored as follows (clockwise from the open end on the left): putative clade IIIb (dark purple), clade II (red), clade IV (pink), clade Vb (light green), clade Va (dark green), clade IIIa (light purple), clade Ia (light blue), and clade Ib (dark blue). Sequences of proteins with known functions are shaded in light blue; BOS is marked by a red arrow and shaded in yellow. BAHD, Benzylalcohol O-acetyltransferase, Anthocyanin O-hydroxycinnamoyltransferase, anthranilate N-Hydroxycinnamoyl/benzoyltransferase, Deacetylvindoline 4-O-acetyltransferase.