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In Hap-W box populations of dry areas, the expression of RAP2.12 is enhanced by other WRKY 12/33 or other unknown genes(s) through the W box cis-element to promote survival under drought stress. In Hap-WT box populations, the shift of the RAP2.12 regulatory cis-element from W to WT box allows WRKY70 to enhance its expression through WT box and promote the worldwide humid colonization in A. thaliana.
