TABLE 4.
Pairwise amino acid and nucleotide sequence analysis of FrxA proteins from 11 pairs of metronidazole-sensitive and -resistant H. pylori isolates
H. pylori strain | MIC (μg/ml) | Genetic alterationsa in:
|
|
---|---|---|---|
Amino acid sequenceb | Nucleotide sequencec | ||
5413 | 8 | R at position 206 to H | Missense mutation (G to A at position 617)d |
6102 | 8 | W at position 137 to R; E at position 164 to G | Missense mutations (T to A at position 409; A to G at position 491)d |
1838 | 32 | Generated a stop codon at position 46 | Frameshift due to one nucleotide (G) deletion at position 46 |
6413 | 32 | C at position 161 to Y | Missense mutation (G to A at position 482) |
1950 | 64 | Generated a stop codon at position 74 | Frameshift due to one nucleotide (G) deletion at position 208 |
2312 | 64 | W at position 49 to stop codon | Nonsense mutation (G to A at position 146) |
2228 | 128 | Generated a stop codon at position 74 | Frameshift due to one nucleotide (G) deletion at position 208 |
1740 | 128 | Generated a stop codon at position 74 | Frameshift due to one nucleotide (G) deletion at position 208 |
1857 | 128 | Q at position 168 to stop codon | Nonsense mutation (C to T at position 505) |
5078 | 128 | Generated stop codons at positions 22 and 126 | Frameshift due to one nucleotide (A) addition at position 55 and one nucleotide (A) deletion at position 316 |
8076 | 128 | Generated a stop codon at position 74 | Frameshift due to one nucleotide (G) deletion at position 208 |
Pairwise amino acid and nucleotide sequence alignment was performed against the metronidazole-sensitive partner strain from the same patient who failed anti-H. pylori therapy containing metronidazole. If the FrxA proteins from metronidazole-resistant strains were not truncated, the amino acid sequences were aligned not only with the partner metronidazole-sensitive strain but also with all other metronidazole-sensitive FrxA amino acid sequences, including H. pylori ATCC 700392 (26) and H. pylori J99 (1).
The initiation amino acid (methionine) was counted as +1.
Nucleotide A from the first codon (ATG) for FrxA was counted as +1.