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. 2022 May 2;10:e13366. doi: 10.7717/peerj.13366

Table 1. Ethogram of behaviors used to construct behavioral budgets for coyotes.

Observers coded behavior based on the behavior and descriptions. Behaviors were then condensed into five broad categories for analysis to reduce the number of behaviors and potential for interobserver error. Modified from Shivik et al. (2009).

Behavior Description Broad category
Resting Coyote laying or sitting down Resting
Locomotion Purposeful walking, trotting, or running with head up Active
Standing Standing still with head raised NA
Foraging Orientating, stalking, and searching at a slow pace with head lowered Feeding
Eating Coyote eating with visible jaw movement Feeding
Aggressive Teeth bared, biting, growling, chasing mate away from food. Social
Play Playful behavior with mate, tail wagging, non-threatening posture Social
Neutral Social Howling and other behaviors directed at mate that are neither aggressive, nor playful Social
Stereotyping Repetitive movement with no apparent goal that is repeated for greater than 2 cycles. Active
Investigating car* Head and gaze oriented towards prey model from a distance while standing or walking slowly. Feeding/Car
Interacting with car* Behaviors directed towards prey model such as chasing, pouncing, scratching, or biting. Feeding/Car
Scent marking Urinating or defecating –point event NA

Notes.

*

These behaviors were only relevant during car feeding observations.