Figure 5.
The surplus sharing fluxes and
form a closed cycle in the root–shoot model. Blocked connector endings indicate a negative effect. Boxes indicate direct algebraic equations; green circles indicate fast state variables that buffer inputs and outputs. Both fluxes have a (negative) effect on the other flux. The more nitrogen the root shares, the more the shoot can grow, which in turn shares less carbon because its surplus is smaller. In the same way, a bigger surplus of carbon results in a smaller surplus of nitrogen. Top row: all interactions are direct, the model is not uniquely defined. All other rows introduce state variables that define the model uniquely. Middle row: state variables for
and
. Bottom row: state variables for
only (left) and for
only (right).