Table 1.

Classification of Brutus’s drumming communication

Category	Number of drumming events	Number of trees	Observations	Proposed semantic content
2A	2	1	8	resting period initiation
1A + 1B	2	2	8	travel direction change
3A†	3	1	None	–
1A + 2B or 2A + 1B	3	2	6	resting period initiation and travel direction change
4A	4	1	1	–
2A + 2B† or 1A + 3B† or 3A + 1B†	4	2	None	–

Modified from Boesch (1991)