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. 2022 May 26;13:864902. doi: 10.3389/fpsyt.2022.864902

Table 3.

Neuroimaging methods of reviewed studies.

Glutamatergic compound References Imaging sample size (MDD, HC) Imaging technique ROIs for activity/cerebral blood flow/entropy/receptor binding/glucose metabolism measurement Methods for connectivity measurement Timepoint(s) of MDD imaging relative to treatment
Ketamine Abdallah et al. (131) 18, 25 fMRI N/A Average correlation between BOLD time series of a voxel and all other gray matter voxels in brain (GBCr) Pre-treatment and 24 h post-treatment
Abdallah et al. (132) 22 (Cohort A only), 29 HC in Cohort A and 18 HC in Cohort B fMRI N/A Average correlation between BOLD time series of a voxel and all other gray matter voxels in brain (GBCr) • Cohort A: pre-treatment and 24 h post-treatment

• Cohort B: pre-treatment and repeated during each treatment session
Abdallah et al. (133) 56, 0 fMRI N/A Average correlation between BOLD time series of a voxel and all other gray matter voxels in brain (GBCr) Pre-treatment, during infusion, and 24 h post-treatment
Carlson et al. (134) 20, 0 18F-FDG-PET • Whole-brain CMRGlu

• ROIs in AMYG, sgACC, anterior HPC, and medial TH
N/A Pre-treatment and ~120 min post-treatment
Chen et al. (135) 48, 0 fMRI N/A • Six ROIs in prefrontal regions (dACC, DLPFC, and mPFC)

• Seeds defined as structures with a 4 mm radius around the coordinates

• FC map of ROIs identified by correlating low-frequency fMRI fluctuations with the seeds
Pre-treatment and on third day after treatment (~48 h post-treatment)
Chen et al. (136) 48, 48 fMRI N/A • Four striatal seeds (limbic, executive, rostral-motor, caudal motor)

• FC map of striatum identified by correlating low-frequency fMRI fluctuations with the ROIs
Pre-treatment
Downey et al. (137) 56, 0 fMRI ROI in sgACC (x = 0, y = 26, z = −8 mm) N/A Pre-treatment and 1 h scanning session which treatment was administered during
Evans et al. (138) 33, 25 fMRI N/A • DMN defined using seed-based correlation method (3dTcorr)

• Average time course from 6 mm radius sphere at PCC (3dROIstats) at the MNI template coordinates of (0, 252, 27) correlated with all other brain voxels
Pre-treatment and ~2 and 10 days after both infusions
Gärtner et al. (139) 24, 0 fMRI N/A • ROIs (MNI): sgACC (sphere at 2, 28, −5 with 5 mm radius), the left and right DLPFC (sphere at ±40, 36, 32 with 10 mm radius), the left and right PCC (sphere at ±6, −50, 24 with 7 mm radius), and the left and right HPC (anatomical seeds from the FSL Harvard-Oxford Atlas)

• Seed-to-voxel correlation maps calculated by extracting residual BOLD time course from seed and computing correlation between that time course and time course of all other voxels; GLM
Pre-treatment and 24 h post-treatment
Gilbert et al. (140) 29, 0 MEG ROIs in left AI (Talairach coordinates: −32, 13, −8) and ACC (Talairach coordinates: −2, 43, 2) N/A Pre-treatment and 6–9 h post-treatment
Kraus et al. (141) 28, 22 fMRI N/A • GBC map transformed to MNI; maps resampled to final voxel size of 3.5 mm isotropic (voxel volume: 42.875 mm3)

• Intra-PFC GBC calculated using same procedures, but GBC calculations restricted to the PFC with a PFC GM mask
Pre-treatment and day 2 or 3 after ketamine or placebo
Lally et al. (142) 20, 0 18F-FDG-PET Whole-brain CMRGlu and ROIs in ventral striatum and OFC N/A Pre-treatment and 2 h post-treatment
Li et al. (143) 48, 0 18F-FDG-PET • ROIs in AMYG and PFC

• Whole-brain voxel-wise analyses
N/A Pre-treatment and immediately after treatment
Loureiro et al. (144) 44, 31 fMRI Whole-brain analysis and ROI in AMYG N/A Pre-treatment and 24–72 h after last infusion
Loureiro et al. (145) 46, 32 fMRI • Cluster in CBM in lobule-VIIb in dorsal-attention part of CBM defined as psychophysiological-interaction seed

• ROIs for CEN (lateral OFC, inferior frontal cortex, supramarginal gyrus, and DLPFC), SN (AI, dACC, and middle frontal cortex) and SMN (posterior INS, precentral and the superior paracentral)
Psychophysiological-interaction analysis using CBM seed generated from the average NoGo-Go activation; GLM Pre-treatment, 24 h after first infusion, and 24 or 72 h after last infusion
McMillan et al. (146) 26, 0 fMRI and EEG sgACC N/A Treatment administered 7 min into a 16 min scan
Mkrtchian et al. (147) 27, 19 fMRI N/A • Seed-to-whole-brain analysis with four striatal seeds (3.5 mm radius spheres): ventral striatum (±9, 9, −8), dorsal caudate (±13, 15, 9), dorsal caudal Pu (±28, 1, 3), and ventral rostral Pu (±20, 12, −3)

• Primary visual cortex used as control region for sensitivity analysis examining whether results were specific to PFC regions or due to global pattern
2 days post-treatment
Morris et al. (148) 42, 20 fMRI ROI in sgACC Computed between sgACC and HPC ROI's using Pearson's correlation Pre-treatment and within 5 days after infusion
Murrough et al. (149) 18, 20 fMRI • Whole-brain, voxel-wise GLM

• Single-subject whole-brain maps reflecting BOLD signals
Investigated FC of regions that demonstrated brain activation main effects; GLM Pre-treatment and 24 h post-treatment
Nakamura et al. (150) 15, 0 fMRI N/A • Whole brain FC maps with seed regions for each hemispheric AMYG and bilateral ventral PCu in the MNI space

• FC between seed and each voxel of whole brain computed as contrast of parameter estimates of the GLM
Pre-treatment and 6–24 h after last infusion
Nemati et al. (151) 258 total (randomized participants were scanned), 0 fMRI N/A GBC computed with average correlation of each voxel/vertex with all other gray matter voxels and vertices Pre-treatment and during infusion
Nugent et al. (152) 13, 18 MEG and 18F-FDG-PET N/A • Data from subjects entered into group independent components analysis to extract 25 components; linear regression used to obtain independent component maps

• ROIs in left and right AMYG and sgACC
Pre-treatment, MEG 6–7 h post-treatment, 18F-FDG-PET ~120 min post-treatment
Reed et al. (153) 33, 26 fMRI Whole brain analysis N/A Pre-treatment and 1–3 days after each infusion
Reed et al. (154) 33, 24 fMRI Whole brain analysis N/A Pre-treatment and 1–3 days after each infusion
Roy et al. (155) 11, 0 fMRI • 132 ROIs

• Cortical and subcortical from the FSL Harvard-Oxford atlas

• Cerebellar regions from the automated anatomical labeling atlas
N/A Pre-treatment and 1 day after last infusion
Sahib et al. (156) 22, 18 Perfusion MRI Regional cerebral blood flow values from ROIs that showed significant changes in whole brain analysis N/A Pre-treatment, 24 h after first infusion, 24–72 h after last infusion
Sahib et al. (157) 47, 32 fMRI • Whole brain analysis

• ROIs in right and left SMA
N/A Pre-treatment, 24 h after first infusion, 24–72 h after last infusion
Sahib et al. (158) 61, 40 fMRI N/A • Nodes generated with high-dimensional group independent components analysis

• Network modeling performed using partial temporal correlation between node time series
Pre-treatment, 24 h after first infusion, 24–72 h after last infusion
Salvadore et al. (159) 11, 11 MEG • False discovery rate calculated using a ROI approach encompassing the ACC/Brodmann area 24/32

• AMYG exploratory analyses
N/A Pre-treatment
Salvadore et al. (160) 15, 0 MEG • False discovery rates estimated within ROI encompassing the ACC/Brodmann area 24/32 • Dynamic imaging of coherent sources using single seed point in the pgACC

• ROI encompassing bilateral AMYG
Pre-treatment
Siegel et al. (161) 23, 27 fMRI N/A Exploratory FC analysis to visualize connectivity between three targets (DMN, sgACC, limbic system) and the rest of the brain Pre-treatment and 2 weeks post-treatment
Sterpenich et al. (162) 10, 0 fMRI • Whole brain analysis with anatomical masks for AMYG, INS, ACC, OFC, and ventral striatum

• ROIs in medial substantia nigra/VTA
N/A Pre-treatment and 1 and 7 days post-treatment
Thai et al. (163) 11, 0 fMRI ROIs in left and right HPC, left and right AMYG, subcallosal cortex, ACC, left and right NAc, PCu, and PCC N/A Pre-treatment and 1 day following last infusion
Tiger et al. (164) 30, 0 PET ROI in CBM N/A Pre-treatment and 24–72 h post-treatment
Vasavada et al. (165) 44, 50 fMRI N/A • DMN, CEN, and SN chosen to investigate FC with HPC (right and left) and AMYG (right and left)

• Time courses for AMYG and HPC extracted ROI masks derived from the Harvard–Oxford subcortical structural atlases

• Correlations calculated between time courses of the networks and seeds
Pre-treatment, 24 h after first infusion, 24–72 h after last infusion
Wang et al. (166) 60 (MDD and bipolar without psychotic symptoms), 0 fMRI N/A • Left and right habenula identified with centers of MNI coordinates (−2.8, −24.2, 2.3) and (4.8, −24.1, 2.2), respectively

• Both of which have a radius of 3 mm
Pre-treatment
Memantine Kilpatrick et al. (167) 26, 0 fMRI N/A Anterior and posterior DMN nodes (PCu and mPFCs) with dual regression to create individual DMN parameter estimate maps Pre-treatment and 3-month follow-up
D-cycloserine Cole et al. (168) 10, 12 EMG M1 localized using neuronavigation and electromyographic electrodes positioned over right first dorsal interosseous muscle N/A 1–2 h post-treatment
Nitrous oxide
4-chlorokynurenine Park et al. (169) 12, 0 1H-MRS and fMRI pgACC 30 ROIs previously identified as being associated with MDD using a 6 mm sphere Pre-treatment and ~60–120 min post-treatment
Lanicemine Abdallah et al. (133) 56, 0 fMRI N/A Average correlation between BOLD time series of a voxel and all other gray matter voxels in brain (GBCr) Pre-treatment, during infusion, and 24 h post-treatment
Downey et al. (137) 56, 0 fMRI ROI in sgACC (x = 0, y = 26, z = −8 mm) N/A Pre-treatment and 1 h scanning session which treatment was administered during

1H-MRS, proton magnetic resonance spectroscopy; 18F-FDG, fluorodeoxyglucose; AI, anterior insula; ACC, anterior cingulate cortex; AMYG, amygdala; BOLD, blood-oxygen-level-dependent; CBM, cerebellum; CEN, central executive network; CMRGlu, cerebral metabolic rate for glucose utilization; CN, caudate nucleus; dACC, dorsal anterior cingulate cortex; DLPFC, dorsolateral prefrontal cortex; DMN, default mode network; EEG, electroencephalogram; EMG, electromyography; FC, functional connectivity; fMRI, functional magnetic resonance imaging; GBC, global brain connectivity; GBCr, global brain connectivity with global signal regression; GLM, general linear modeling; GM, gray matter; HC, healthy control subjects; HPC, hippocampus; INS, insula; MDD, major depressive disorder participants; MEG, magnetoencephalography; MNI, Montreal Neurological Institute; mPFC, medial prefrontal cortex; MRI, magnetic resonance imaging; N/A, not applicable/available; NAc, nucleus accumbens; OFC, orbitofrontal cortex; PCC, posterior cingulate cortex; PCu, precuneus; PET, positron emission tomography; PFC, prefrontal cortex; pgACC, pregenual anterior cingulate cortex; Pu, putamen, ROI, region of interest; sgACC, subgenual anterior cingulate cortex; SMA, supplementary motor area; SN, salience network; TH, thalamus; VTA, ventral tegmental area.