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. 2022 Jun 22;21:133. doi: 10.1186/s12943-022-01596-8

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© The Author(s) 2022

Open AccessThis article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated in a credit line to the data.

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Fig. 3 — UCOGCP held distinct ductal profile. A, Major clusters of the ductal type I cells shown in UMAP. B, Top three markers of each cluster obtained from “FindAllMarkers” function from Seurat package (4.0.4) shown in dot plot. C-E, The distribution of each cluster in each sample shown in UMAP, balloon plot and heatmap, respectively. F, Violin plot showing markers of epithelial cells, cancer stem cells, and epithelial-mesenchymal transition (EMT) cells. G, Hallmarks among clusters shown in dot plot. H, Expression levels of enriched genes in HALLMAKS-MYC-TARGET-V1. I, Survival analysis of gene signatures in H in pancreatic cancer using TCGA-PAAD on website Gepia2 (http://gepia2.cancer-pku.cn/#index). J and K, GSEGO analysis of gene markers in cluster 2. L, Expression level of gene signatures enriched in at least eight of the top GSEGO clusters of cluster 3. M, Survival analysis of gene signatures in L in pancreatic cancer using TCGA-PAAD on website Gepia2 (http://gepia2.cancer-pku.cn/#index). N, Expression level and survival analysis of representative genes in L in pancreatic cancer using TCGA-PAAD on website Gepia (http://gepia.cancer-pku.cn/index.html)