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. 2022 Jun 25;305:135441. doi: 10.1016/j.chemosphere.2022.135441

Table 1.

Classification of water and wastewater viruses as well as virus abundance and removal efficiency in wastewater treatment plant (WWTP).

Sample/Country Treatment technology Virus name Gene Virus abundance (i) genes copies/L
(ii) Gene equivalents/L
Log Removal Values
Quantitative analysis method References
Influent Effluent LRVs
WWTP/Grid, Italy Separation, primary sedimentation, secondary biological treatment and disinfection Enteroviruses s-s RNA 3.3 × 107 (i) 7.6 × 106 (i) 0.63 RT-PCR, Real-time qPCR (La Rosa et al., 2010)
WWTP/Southern Arizona,
USA
Plant A: conventional activated sludge process
Plant B: biological trickling filter process
Pepper mild mottle virus s-s RNA 3.7–4.4 × 
106/3.2–9.4
 × 106 (i)
4.6–6.3 × 105 (i) 0.76–0.99/1.8
±0.2
TaqMan-based qPCR (Kitajima et al., 2014)
WWTP/Arizona, USA Activated sludge and trickling
filter
Coxsackieviruses 3.24 × 105 (i) 1.54 × 103 (i) 2.32 RT-PCR (Kitajima et al., 2015)
WWTP/France Primary decantation and biological secondary treatment Astroviruses s-s RNA 2.69 × 103 (i) RT-qPCR (Prevost et al., 2015)
WWTP/Kampala, Uganda Conventional activated sludge method, in summer
2016
Hepatitis A virus s-s RNA 2.01 × 
103–8.39 × 103 (i)
1.93 × 
103–8.70 × 
103 (i)
qPCR and quantitative
RT-PCR
(O'Brien et al., 2017)
Eastern Cape, South Africa Activated sludge system with 40,000 m3/day flow rate Rotaviruses d-s RNA 1.2 × 105 (i) 2.6 × 104 (i) 0.66 Quantitative
TaqMan real-time PCR
(Osuolale and Okoh, 2017)
WWTP/North Wales, UK With filter beds for secondary treatment and serves ca. 4000 inhabitants Norovirus
Genotypes
GI/GII
s-s RNA 8.8 × 104 (i) 3 × 104 (i) 0.47 RT-qPCR (Farkas et al., 2018)
WWTP/Greater Cairo, Egypt Activated sludge as secondary treatment process with 600,000 m3/day Polyomaviruses d-s DNA 3.9 × 105 (i) 4.51 × 103 (i) 1.93 Real time PCR (Hamza and Hamza, 2018)
Sample/Country Treatment technology Virus name Gene Virus abundance (i) genes copies/L
(ii) Gene equivalents/L
Log Removal Values
Quantitative analysis method References
Influent Effluent LRVs
WWTP
Egypt
330,000 m3/day capacity Adenoviruses d-s DNA 4.3 × 105–8.7 × 106 (i) 1.22 × 104–3.7
 × 106 (i)
Real time PCR Elmahdy et al. (2019)
WWTP
Japan
Conventional activated sludge process SARS-CoV-2 s-s RNA 2.4 × 103 (i) RT-qPCR Haramoto et al. (2018)
WWTP/P. Alegre, Brazil An anaerobic stage by the UASB reactor and an aerobic stage through the Unitank® System. SARS-CoV-2 s-s RNA 4.14 × 101–5.23 × 103 (i) RT-qPCR Zaneti et al. (2021)
WWTP/Murica, Spain Untreated wastewater samples SARS-CoV-2 5.4 ± 0.2 log10 (i) RT-qPCR Zaneti et al. (2021)
Raw WWTP/Paris/France Several major wastewater treatment plants SARS-CoV-2 s-s RNA 3–50 × 103 (ii) RT-qPCR Wurtzer et al. (2020)
Raw wastewaters/Germany Untreated sewage and treated water by ozonation samples SARS-CoV-2 Solid phase 25 (i)
Aqueous phase 1.8 (i)
Solid phase 13 (ii)
Aqueous phase 8.8 (ii)
RT-qPCR Westhaus et al. (2021)

Abbreviations: qPCR: Quantitative polymerase chain reaction, RT-(q)PCR: Reverse Transcriptase-(Quantitative) polymerase chain reaction, ICC-qPCR: Integrated cell culture with quantitative polymerase chain reaction.