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. Author manuscript; available in PMC: 2022 Jul 10.
Published in final edited form as: Theor Popul Biol. 2020 Feb 3;132:24–32. doi: 10.1016/j.tpb.2020.01.005
J11=((α+θ1+μ1+ρ)000r12θ1(α+θ2+μ2+ρ)0000r(n1)nθn1(α+θn+μn+ρ))
J12=(a1β1x1c11a1β1x1c12a1β1x1c1na2β2x2c21a2β2x2c22a2β2x2c2nanβnxncn1anβnxncn2anβnxncnn)
J21=αIn×n,and
J22=((γ+θ1+μ1+ρ)000r12θ1(γ+θ2+μ2+ρ)0000r(n1)nθn1(γ+θn+μn+ρ))