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Total nutrient content in plant tissues. Total carbon (C), nitrogen (N), sulfur (S), phosphorus (P), and potassium (K) in standing aboveground biomass, belowground biomass, photosynthetic tissue, and annual litter inputs at each thaw stage (palsa, bog, fen). Letters indicate significant differences between different identified thaw stages (P, palsa; B, bog; F, fen) in total pool size or flux (ANOVA, Tukey HSD p < .05). Error bars are the standard error of the mean
