Table 2.
Model averaged parameter estimates (Est.), standard errors (SE), and relative variable importance (RVI) of (1) deadwood and forest characteristics, (2) microclimatic variables, and (3) combination of deadwood and forest characteristics, and microclimatic variables, explaining (a) current and (b) long-term occurrence, and (c) current and (d) long-term abundance, based on models with ΔAICc < 7
| Variables | Est | SE | RVI | Est | SE | RVI | Est | SE | RVI | Est | SE | RVI | ||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1. Deadwood and forest characteristics | (a) Current occurrence | (b) Long-term occurrence | (c) Current abundance | (d) Long-term abundance | ||||||||||
| Diameter | 0.279 | 0.085 | 0.99 | 0.254 | 0.084 | 1.00 | 0.048 | 0.039 | 0.70 | 0.045 | 0.014 | 1.00 | ||
| Length | < 0.001 | < 0.001 | < 0.01 | < 0.001 | < 0.001 | 0.56 | ||||||||
| Ground contact | < 0.001 | 0.004 | 0.17 | 0.001 | 0.005 | 0.19 | − 0.001 | 0.002 | 0.19 | |||||
| Bark cover | 0.005 | 0.013 | 0.24 | 0.001 | 0.009 | 0.19 | − 0.001 | 0.006 | 0.15 | |||||
| Vegetation cover | 0.002 | 0.009 | 0.18 | − 0.001 | 0.010 | 0.18 | < 0.001 | 0.003 | 0.14 | − 0.006 | 0.005 | 0.62 | ||
| Softness | − 0.079 | 0.134 | 0.39 | 0.003 | 0.062 | 0.18 | 0.072 | 0.065 | 0.68 | 0.120 | 0.022 | 1.00 | ||
| Canopy openness | 0.002 | 0.011 | 0.02 | 0.001 | 0.004 | 0.15 | 0.028 | 0.004 | 1.00 | |||||
| Basal area | − 0.173 | 0.061 | 0.98 | − 0.228 | 0.064 | 0.99 | 0.002 | 0.012 | 0.14 | |||||
| Vegetation type | − 0.217 | 0.564 | 0.26 | − 0.597 | 0.894 | 0.45 | 0.115 | 0.223 | 0.32 | − 0.149 | 0.207 | 0.38 | ||
| Stand type 2 | − 0.002 | 0.097 | < 0.01 | − 0.004 | 0.081 | 0.10 | ||||||||
| Stand type 3 | 0.008 | 0.226 | − 0.208 | 0.709 | ||||||||||
| Stand type 4 | − 0.028 | 0.346 | − 0.055 | 0.232 | ||||||||||
| 2. Microclimatic variables | (a) Current occurrance | (b) Long-term occurrance | (c) Current abundance | (d) Long-term abundance | ||||||||||
| Average temperature | ||||||||||||||
| (A) Autumn | − 0.428 | 0.899 | 0.27 | 0.067 | 0.391 | 0.04 | − 0.053 | 0.181 | 0.14 | 0.003 | 0.028 | 0.03 | ||
| (B) Winter | − 0.085 | 0.411 | 0.05 | − 2.616 | 1.967 | 0.71 | − 0.046 | 0.175 | 0.16 | − 0.664 | 0.186 | 0.95 | ||
| (c) Spring | 0.065 | 0.192 | 0.15 | 0.627 | 0.619 | 0.62 | − 0.007 | 0.034 | 0.10 | 0.047 | 0.058 | 0.46 | ||
| (d) Summer | 0.164 | 0.326 | 0.26 | 0.240 | 0.471 | 0.26 | − 0.029 | 0.076 | 0.17 | 0.013 | 0.037 | 0.14 | ||
| Temperature fluctuations | ||||||||||||||
| (a) Autumn | − 0.030 | 0.335 | 0.16 | − 0.013 | 0.399 | 0.18 | − 0.037 | 0.116 | 0.15 | 0.005 | 0.026 | 0.05 | ||
| (b) Winter | − 2.318 | 1.013 | 0.94 | − 0.687 | 1.181 | 0.29 | − 0.088 | 0.206 | 0.26 | − 0.016 | 0.078 | 0.07 | ||
| (c) Spring | 0.018 | 0.079 | 0.09 | 0.002 | 0.031 | 0.01 | − 0.004 | 0.021 | 0.08 | 0.001 | 0.005 | 0.03 | ||
| (d) Summer | 0.010 | 0.052 | 0.08 | − 0.029 | 0.097 | 0.16 | − 0.003 | 0.018 | 0.08 | 0.001 | 0.007 | 0.05 | ||
| Temperature extremes | ||||||||||||||
| (a) Winter | − 0.075 | 0.349 | 0.13 | − 0.551 | 1.255 | 0.20 | − 0.016 | 0.182 | 0.12 | − 0.032 | 0.145 | 0.05 | ||
| (b) Summer | 0.062 | 0.146 | 0.20 | 0.004 | 0.045 | 0.02 | − 0.016 | 0.041 | 0.18 | 0.003 | 0.011 | 0.07 | ||
| Moisture | ||||||||||||||
| (a) Autumn | − 0.002 | 0.016 | 0.12 | 0.003 | 0.027 | 0.13 | − 0.001 | 0.005 | 0.09 | − 0.026 | 0.006 | 1.00 | ||
| (b) Winter | 0.006 | 0.013 | 0.30 | − 0.007 | 0.015 | 0.30 | 0.003 | 0.007 | 0.28 | 0.015 | 0.004 | 1.00 | ||
| (c) Spring | < 0.001 | 0.004 | 0.10 | − 0.001 | 0.006 | 0.12 | 0.003 | 0.006 | 0.25 | |||||
| (d) Summer | − 0.007 | 0.015 | 0.31 | 0.002 | 0.010 | 0.15 | 0.002 | 0.005 | 0.26 | |||||
| 3. All variables combined | (a) Current occurrence | (b) Long-term occurrence | (c) Current abundance | (d) Long-term abundance | ||||||||||
| Diameter | 0.274 | 0.093 | 1.00 | 0.281 | 0.118 | 1.00 | 0.040 | 0.067 | 0.33 | 0.109 | 0.028 | 0.97 | ||
| Length | 0.001 | < 0.001 | 0.72 | |||||||||||
| Ground contact | < 0.001 | 0.003 | 0.06 | 0.001 | 0.005 | 0.07 | < 0.001 | 0.001 | 0.03 | |||||
| Bark cover | 0.001 | 0.006 | 0.09 | < 0.001 | 0.004 | 0.05 | < 0.001 | 0.001 | 0.01 | |||||
| Vegetation cover | 0.001 | 0.007 | 0.09 | 0.005 | 0.020 | 0.10 | − 0.003 | 0.008 | 0.12 | |||||
| Softness | − 0.038 | 0.104 | 0.18 | − 0.011 | 0.072 | 0.06 | 0.161 | 0.096 | 0.81 | 0.010 | 0.028 | 0.13 | ||
| Canopy openness | 0.002 | 0.011 | 0.04 | 0.002 | 0.013 | 0.02 | < 0.001 | 0.001 | 0.02 | 0.015 | 0.010 | 0.75 | ||
| Basal area | − 0.025 | 0.055 | 0.21 | − 0.051 | 0.089 | 0.27 | 0.023 | 0.049 | 0.22 | 0.006 | 0.018 | 0.12 | ||
| Vegetation type | − 0.156 | 0.544 | 0.13 | − 0.893 | 1.582 | 0.32 | 0.033 | 0.149 | 0.07 | |||||
| Stand type 2 | 0.003 | 0.896 | 0.21 | |||||||||||
| Stand type 3 | − 2.965 | 1108 | ||||||||||||
| Stand type 4 | − 0.958 | 2.053 | ||||||||||||
| Average temperature | ||||||||||||||
| (a) Autumn | − 0.074 | 0.466 | 0.09 | 0.099 | 0.715 | 0.07 | 0.005 | 0.096 | 0.021 | 1.204 | 0.435 | 0.92 | ||
| (b) Winter | − 0.757 | 1.095 | 0.39 | − 0.911 | 1.563 | 0.30 | − 0.265 | 0.516 | 0.27 | − 0.065 | 0.229 | 0.08 | ||
| (c) Spring | 0.011 | 0.086 | 0.05 | 0.068 | 0.282 | 0.10 | − 0.002 | 0.019 | 0.03 | |||||
| (d) Summer | 0.203 | 0.415 | 0.24 | 0.251 | 0.639 | 0.17 | − 0.002 | 0.023 | 0.03 | |||||
| Temperature fluctuations | ||||||||||||||
| (a) Autumn | 0.038 | 0.254 | 0.06 | 0.205 | 0.742 | 0.12 | < 0.001 | 0.022 | 0.02 | |||||
| (b) Winter | − 0.450 | 0.857 | 0.28 | − 0.230 | 0.816 | 0.12 | − 0.013 | 0.085 | 0.04 | |||||
| (c) Spring | 0.006 | 0.048 | 0.04 | 0.012 | 0.076 | 0.06 | 0.002 | 0.014 | 0.03 | |||||
| (d) Summer | 0.032 | 0.112 | 0.11 | 0.019 | 0.088 | 0.07 | 0.001 | 0.008 | 0.02 | |||||
| Temperature extremes | ||||||||||||||
| (a) Winter | − 0.089 | 0.436 | 0.06 | − 0.738 | 1.761 | 0.17 | − 0.436 | 0.845 | 0.27 | − 1.777 | 0.577 | 0.92 | ||
| (b) Summer | 0.112 | 0.218 | 0.27 | 0.026 | 0.112 | 0.08 | < 0.001 | 0.006 | 0.01 | 0.006 | 0.028 | 0.05 | ||
| Moisture | ||||||||||||||
| (a) Autumn | < 0.001 | 0.012 | 0.07 | 0.001 | 0.017 | 0.05 | < 0.001 | 0.002 | < 0.01 | − 0.001 | 0.005 | 0.03 | ||
| (B) Winter | < 0.001 | 0.005 | 0.07 | − 0.013 | 0.025 | 0.28 | 0.001 | 0.005 | 0.11 | 0.001 | 0.003 | 0.03 | ||
| (C) Spring | − 0.001 | 0.006 | 0.07 | − 0.002 | 0.008 | 0.08 | 0.004 | 0.008 | 0.24 | |||||
| (d) Summer | − 0.002 | 0.008 | 0.11 | < 0.001 | 0.004 | 0.04 | 0.007 | 0.008 | 0.47 | |||||
For stand and vegetation types, the first categories are taken as references. Variable importance > 0.5 is highlighted in bold font