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. 2022 Jul 26;13:4314. doi: 10.1038/s41467-022-31873-5

Table 1.

Summary of the source GWAS and number of causal proteins for each brain trait identified through PWAS, SMR/HEIDI, and COLOC

GWAS statistics Causal protein counts Causal proteins shared among traits
N significant SNPs cis trans total Psychiatric Neuro-degenerative Brain structure
Psychiatric
Major depressive disorder 807,553 9744 96 2 98 46% 2% 8%
Bipolar disorder 51,710 240 70 0 70 44% 3% 7%
Schizophrenia 77,096 14,724 156 2 158 31% 3% 7%
Anxiety 175,163 166 4 0 4 75% 50% 25%
Post-traumatic stress disorder 214,408 4607 16 0 16 44% 6% 13%
Alcoholism 300,789 1857 19 1 20 45% 10% 5%
Neuroticism 390,278 7759 122 2 124 34% 4% 6%
Insomnia 386,533 463 29 0 29 41% 0% 10%
Neurodegenerative
Alzheimer’s disease (AD1) 455,258 2357 16 2 18 39% 22% 6%
Alzheimer’s disease (AD2) 63,926 1506 3 1 4 50% 50% 0%
Frontotemporal dementia 12,908 29 0 0 0
Amyotrophic lateral sclerosis 80,610 182 6 0 6 17% 0% 0%
Lewy body dementia 6618 189 0 0 0
Parkinson’s disease 482,730 3464 19 1 20 25% 10% 15%
Brain structure
Hippocampal volume 26,814 461 2 0 2 0% 0% 50%
Nucleus accumbens volume 32,562 138 3 0 3 0% 0% 0%
Amygdala volume 34,431 0 0 0 0
Putamen volume 37,571 1465 6 0 6 50% 17% 50%
Brainstem volume 28,809 1063 10 0 10 50% 10% 30%
Caudate nucleus volume 37,741 141 7 0 7 14% 0% 29%
Globus pallidus volume 34,413 272 7 0 7 14% 14% 43%
Thalamus volume 34,464 50 2 0 2 50% 0% 50%
Cortical thickness 33,992 370 19 0 19 37% 5% 21%
White matter hyperintensity 48,454 1340 19 1 20 30% 0% 15%
Cortical surface area 33,992 1127 20 0 20 40% 15% 15%

The sample size (N) for each trait is the number of samples that contributed to the GWAS results used in our analysis, which may be smaller than the sample reported in the original GWAS publication due to data access limitations or restricting to samples of European ancestry. GWAS-significant SNPs were defined as SNPs with p value <5 × 10−8. A protein is considered shared with another trait group if it is also causal for any other trait in that group. Since different traits have different number of identified causal proteins, we examined the percentage of the causal proteins of each trait that is shared within a trait group or between two trait groups. This information is also depicted in Figs. 3A and 4A.