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. 2022 Jul 19;16:893587. doi: 10.3389/fnins.2022.893587

TABLE 3.

Summary of characterizing flavors (sans menthol) on nicotine use.

Flavor Species Route Concentration/Dose Nicotine pH Main findings References
Sucrose Wistar rats, adult males Oral Sucrose (0–10%) n/a • Rats increase consumption of nicotine when solutions are sweetened with sucrose Smith and Roberts, 1995
Nicotine (10 μg/ml)
“Vanilla,” “Coconut” C57BL/6J mice, adult males Oral Vanilla, coconut (0.01–1%)
Nicotine (40–120 μg/ml)
7.7–7.9 • Mice consume more vanilla-flavored (1%) nicotine solutions (60 mg/ml) than nicotine only solutions Tannous et al., 2021
• Vanilla flavor enhances oral nicotine self-administration compared to nicotine alone
“Retro Fruit Twist,” “Tobacco” C57BL/6J mice, adult males Oral Flavors (n/a)
Nicotine (30–200 μg/ml)
Free-base • Mice consume more fruit-flavored nicotine solutions than nicotine-only solutions (75, 100, and 200 mg/ml) Wong et al., 2020
• Mice do not show increased consumption of tobacco-flavored nicotine solutions compared to nicotine-only solutions
“Strawberry” C57BL/6J mice, adolescent-adult males and females Oral Strawberry (Unsweetened Strawberry Kool-Aid made in 2% saccharin) Free-base • Adolescent mice prefer strawberry-flavored nicotine solutions over nicotine-only solutions Patten et al., 2021
Nicotine (0.1 mg/ml) • Adolescent females show greater preference than adolescent males in this effect
“Chocolate,” “Grape” Sprague Dawley rats, adolescent females Oral, intravenous (i.v.) Chocolate (0.5% Hershey’s Unsweetened Cocoa) Free-base • Rats do not self-administer i.v. nicotine with contingent intraoral flavor delivery Chen et al., 2011
Grape (0.1% Unsweetened Grape Kool-Aid)
Nicotine (15–30 μg/kg/infusion)
*0.4% saccharin added to oral solutions
“Licorice” Sprague Dawley rats, adult males Oral, i.v. Licorice (0.1, 1.0% vol/vol licorice root extract) Free-base • Licorice (1.0%) as a conditioned reinforcer prior to self-administration testing increases operating responding for nicotine infusions whereas unconditioned licorice does not Palmatier et al., 2020
Nicotine (7.5 μg/kg/infusion)
Saccharin, sucrose Sprague Dawley rats, adult males Oral, i.v. Saccharin (0.32%) Free-base • Contingent delivery of intraoral sucrose or saccharin enhances self-administration of i.v. nicotine obtained via lever pressing Wickham et al., 2018
Sucrose (10%)
Nicotine (0, 30 μg/kg/infusion)
“Sweet flavors” – peach, watermelon, blackberry, cotton candy, cola, sweet lemon tea Human, N = 20, 55% male, age 19–34) Inhalation Flavors (n/a)
Nicotine (0, 6 mg/ml)
Free-base • Sweet-flavored e-cigarettes increase appeal ratings compared to tobacco, menthol, and unflavored e-cigarettes Goldenson et al., 2016
“Cherry Crush,” “Vivid Vanilla,” “Piña Colada,” “Peach Schnapps” Human, N = 31, 58% male, average age = 34)
*age range n/a
Inhalation Flavor (n/a)
Nicotine (12 mg/ml)
Free-base • Piña Colada rated as sweetest and most liked
• Sweetness is positively associated with liking
• Harshness is negatively associated with liking
Kim et al., 2016
“Cherry,” “Chocolate” Human, N = 132, 49% male, age 18–45) Inhalation Flavors (n/a)
Nicotine (18 mg/ml)
Free-base • Individuals rate cherry and chocolate e-cigarettes as sweeter than unflavored e-cigarettes, but not more liked Mead et al., 2019
• Sweetness is positively associated with liking
• Irritation and bitterness are negatively associated with liking
“Cherry,” “Chocolate” Human, N = 39, 100% male, age 18–45 Inhalation Flavor (n/a)
Nicotine (6, 18 mg/ml)
n/a • Sweetness is positively associated with first puff liking Baker et al., 2021a
• Harshness/irritation is negatively associated with first puff liking
• First puff liking is not associated with total nicotine intake
“Cherry” Human, N = 19, 68% male, age 21–35) Inhalation Cherry (4.7% or 9.3% vol/vol)
Nicotine (0, 6, 12 mg/ml)
Free-base • Increasing nicotine concentration increases ratings of bitterness and reduces appeal Pullicin et al., 2020
• Cherry flavor increases ratings of sweetness and liking
• Increasing the concentration of cherry flavor from 4.7 to 9.3% increases perceived sweetness, harshness, and bitterness but does not alter hedonic ratings