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. 2022 Jul 22;13:954472. doi: 10.3389/fpsyg.2022.954472

TABLE 3.

Ethogram of coded behaviors.

Category Variable Description
Latency choice Latency to consume LVR/HVR Time from start of trial until horse either closed his/her lips around the LVR or until the delay time has passed
Attention Look away Head turned away in >45° angle from experimenter
Distance Large distance Horses’ mouth is more than 0.5 m away from the hand holding the LVR
Other behaviors Empty chewa Horse chews without having food in his/her mouth
Oral manipulation Licking, nibbling or biting into barrier/box/door or own body parts
Head movement Any repeated movements with the head (i.e., horizontal and vertical movements or rotational movements)
Flehmen Lifting the upper lip, usually associated with a forward stretched neck
Pawing Repeatedly lifting one leg and scratching with the hoof on the ground
Reward-directed behaviors Sniffing LVR Sniffing on LVR without taking it into the mouth
Pushing LVR Pushing hand holding LVR away with mouth or head

aOnly coded after 10 s of a trial had elapsed to avoid coding instances of horses still chewing the previous reward.