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. 2022 Jul 28;16:951202. doi: 10.3389/fncel.2022.951202

Table 1.

Summary of the main targets of cdk5 in neurological diseases and mechanism of Cdk5 and nervous system diseases.

Targets or Pathways References
Nervous system diseases Alzheimer’s disease Aβ generation: Cdk5-PPAR-β, Ginsenoside Rg1-Aβ-Cdk5/PPARγ pathway, Cdk5/Calcineurin signaling, Cdk5/Drp1-mediated mitochondrial fission, GSK-3 β and TrkA/Cdk5, Cdk5/HIF-1 (Mawuenyega et al., 2010; Palop and Mucke, 2010; Cheung and Ip, 2012; Jaiswal and Sharma, 2017; Lazarevic et al., 2017; Guo et al., 2018; Fang et al., 2019; Lapresa et al., 2019; Quan et al., 2019, 2020; Ribeiro Filho et al., 2019; Chao et al., 2020; Li et al., 2020a; Park et al., 2020; Ai et al., 2022)
Tau phosphorylation: Leukotriene/Cdk5, Cdk5/mitochondria and Golgi function, Cdk5/MARK4, Cdk5/RPS23RG1, miR-148a-3p/p35/PTEN signaling pathway, Quercetin-Ca2+- calpain-p25-Cdk5 pathway, miR-504–3p and CDK5 axis (Mohamed et al., 2017; Shen et al., 2018; Wilkaniec et al., 2018; Giannopoulos et al., 2019; Saito et al., 2019; Xu et al., 2019; Huang et al., 2020; Zeng et al., 2021; Zhao et al., 2021; Tanaka et al., 2022)
Neurodegenerative diseases Cdk5 inhibitor in AD: Cdk5/Mcl-1, Cdk5-/ALDH1A1, Cdk5/miR-125b, Nano-HO-JNK/Cdk5/GSK-3β, Kaixinsan/ GSK-3β and CDK5 (Nikhil and Shah, 2017; Shukla et al., 2017; Nikhil et al., 2019; Zeb et al., 2019a, b; Zhuang et al., 2020a, b; Qu et al., 2021; Chen et al., 2022; Jiao et al., 2022)
Parkinson’s disease Cdk5/inflammation, Cdk5/immune hyperactivity, Cdk5/mitochondrial fission, Cdk5/degradation of ubiquitin ligases, Cdk5-Luteolin, phosphorylation induced SIRT2 nuclear translocation (He et al., 2018; Wang et al., 2018; Park et al., 2019; Shukla et al., 2019; Cheng et al., 2020; Reudhabibadh et al., 2021; Yan et al., 2022)
Huntington’s disease P25/Cdk5, Cdk5-drp1, Cdk5/DARPP-32 (Paoletti et al., 2008; Langhorne et al., 2011; Cherubini et al., 2015; Brito et al., 2019)
Ischemia Stroke Cdk5/Zinc chelator, Cdk5/neuregulin-1 β, Cdk5/TFP5, Cdk5/tat-Cdk5 CTM, Cdk5/ERK1/2 signaling pathway, Cdk5/inhibition of p53 dependent apoptosis, Cdk5/trkb-erk1/2-creb pathway, Cdk5/phosphorylation of drp1s616, miR-148b-3p via CDK5R1/SIRT1 (Becerra-Calixto and Cardona-Gomez, 2017; Cui et al., 2017; Ji et al., 2017; Liu et al., 2017; Zhao et al., 2017; Munoz-Manco et al., 2018; Tuo et al., 2018; Zhang et al., 2018; Shin et al., 2019; Zhu et al., 2019; Chen et al., 2021, 2022)
Cerebrovascular disease Intracerebral Hemorrhage Cdk5-ATM signalin pathway, Cdk5/P35, Cdk5- p75NTR (Wu et al., 2016; Roufayel and Murshid, 2019; Zhou et al., 2019)
Neuropathic Pain Cdk5/CREB, Cdk5/PPAR γ pathway, mir-196a-5p/Cdk5 axis, Cdk5/CRMP2, Cdk5-NR2A pathway, Cdk5/TRPA1 (Li et al., 2014, 2020; Yang et al., 2014; Chernov et al., 2018; Sulak et al., 2018; Moutal et al., 2019; Zhong et al., 2019; Gomez et al., 2020a, 2021; Zhu et al., 2021)
Epilepsy Cdk5/mitochondrial fragmentation, Cdk5/neuroinflammation, Cdk5/endoplasmic reticulum stress, Cdk5/p38 MAPK mediated microglial response (Tian et al., 2008, 2010; Li et al., 2016; Kim and Kang, 2017, 2018; Liu et al., 2017; Hiragi et al., 2018; Kim et al., 2019; Fan et al., 2020; Lee and Kim, 2021)
Glioblastoma Cdk5/PIKE-A, Cdk5/DRP1, miR-21/Cdk5, Cdk5/TP5, Cdk5/DYRK1A, Cdk5/AC1MYR2, Cdk5/TRIM59, OGT/CDK5/ACSS2 pathway (Liu et al., 2008; Ren et al., 2015; Xie et al., 2015; Gonzalez-Vera et al., 2016; Sang et al., 2019; Peyressatre et al., 2020a, b; Tabouret et al., 2020; Chen et al., 2021; Zhou et al., 2021; Ciraku et al., 2022)
Multiple sclerosis Cdk5/oligodendrocytes, Cdk5/lymphocyte activation (Pareek et al., 2010; Luo et al., 2016, 2018)
Other neurological disorders Cdk5/ERK1/2 pathway, Cdk5/caspase-3 pathway, Cdk5/CRMP-2, Cdk5/mitochondrial kinetic defects, Cdk5/Oxidative stress, Cdk5/endoplasmic reticulum stress, nestin-Cdk5-drp1 (Lindqvist et al., 2017; Guo et al., 2018; Kamiki et al., 2018; Roach et al., 2018; Shi et al., 2018; Spurrier et al., 2018; Wang et al., 2018; Barrett et al., 2019; Chen et al., 2019; Liu et al., 2019; Sase et al., 2019; Li et al., 2020b; Rong et al., 2020; Shukla and Singh, 2020, 2022; Xia et al., 2020; Zhang et al., 2021; Daniels et al., 2022; Manglani and Dey, 2022; Takahashi et al., 2022b; Umfress et al., 2022; Zhou et al., 2022)

Abbreviations: PPAR γ, Peroxisome proliferator-activated receptor gamma; APP, Aβ precursor protein; GSK-3 β, Glycogen Synthase Kinase-3; Drp, dynamin-related protein; MARK4, microtubule-affinityregulating kinase 4; TrkA, tropomyosin-relatedkinaseA; GSK-3 β, Glycogen synthase kinase; HIF-1, Hypoxia-inducible factor 1; RPS23RG1, The type Ib transmembrane protein; Mcl-1, myeloid-cell-leukemia-sequence-1; ALDH1A1, aldehyde dehydrogenase 1 family member A1; Fbxw7, F-box/WDrepeat-containing protein 7; TFP5, a modified truncated 24-aa peptide; DARPP-32, dopamine- and cyclic-AMP-regulated phosphoprotein of molecular weight 32,000; OGDR, The oxygen glucose deprivation reperfusion; MEF2D, myocyte enhancer factor 2D; p75 NTR, p75 neurotrophic factor receptor; ATM, Ataxia Telangiectasia Mutated; ERK1, Extracellular signal-regulated kinase 1; CREB, a transcription factor; TRPA1, transient receptor potential action channel 1; NR2A, N-methyl-D-aspartate receptor subunit 2A; NMDAR, N-methyl-D-aspartate receptor; MAPK, mitogen-activated protein kinases; TP5, a thymopentin; MTLE-HS, mesial temporal lobe epilepsy with hippocampal sclerosis; PIKE-A, Isoform A of phosphatidylinositol 3-kinase enhancer; DYRK1A, Dual-specificity tyrosine phosphorylation-regulated kinase 1A; ACSS2, acetate-dependent acetyl CoA synthetase 2; CRMP2, collapsin response mediator protein 2.