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. 2022 Jul 29;9:904339. doi: 10.3389/fmed.2022.904339

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Copyright © 2022 Schillinger, Mwakamui, Mulenga, Tembo, Hodges, Besa, Chandwe, Owino, Edwards, Kelly and Morrison.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) and the copyright owner(s) are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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Biopsies from adult Zambian study participants (A,B) histology, showing variable villus blunting; (C) morphometry; and (D,E) immunostaining for sucrose-isomaltase, showing lower (D) and higher (E) protein expression.