Table 1. Model selection for the telomere/centromere effect (n = 37 species with a centromere position, 425 chromosomes).

Three competing models were compared based on the adjusted R², p-value and AIC-BIC criteria among chromosomes (the best supported model is in bold characters). The number of species supporting each model was calculated based on the adjusted R² within species, for all species with at least five chromosomes. (1) ‘telomere’ model. (2) ‘telomere + centromere + one CO per arm’ model. (3) ‘telomere + centromere + one CO per chromosome’ model. d(c) is the genetic distance to the centromere. d(1) is the total genetic distance. A second model selection was done on a subset of chromosomes with at least 50 cM on each chromosome arm (n = 36 species, 333 chromosomes).

#	Model	Expected	Adjusted R2	p	AIC	BIC	Species
Full dataset (37 species, 425 chromosomes)
1	Telomere	d(1/2) / d(1) = 0.5	0.22	< 0.001	-477.8	-465.7		0
2	Tel. + Cent. + CO per arm	(d(c)– 50) / (d(1)– 100) = c	-	0.72	3098.2	3110.4		7
3	Tel. + Cent. + CO per chr.	d(c) / d(1) = c	0.51	< 0.001	-476.6	-464.5		30
Subset (36 species, 333 chromosomes)
1	Telomere	d(1/2) / d(1) = 0.5	0.18	< 0.001	-407.5	-396.1		0
2	Tel. + Cent. + CO per arm	(d(c)– 50) / (d(1)– 100) = c	-0.001	0.42	1939.1	1950.5		10
3	Tel. + Cent. + CO per chr.	d(c) / d(1) = c	0.50	< 0.001	-396	-384.6		26