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. 2022 Sep 21;377(1863):20210177. doi: 10.1098/rstb.2021.0177

Table 1.

Overview of muscle activations found in six Carnivora taxa. Caniformia: Czechoslovakian wolfdogs, Bornean sun bears and African wild dogs. Feliformia: spotted hyaenas, meerkatsa and lionsa. Fifty-nine open-mouth faces were coded from Czechoslovakian wolfdogs (Italy), 20 from rehabilitant sun bears at the Bornean Sun Bear Conservation Centre (Malaysia), 10 from wild dogs at the Dvur Kralove Zoo (Czech Republic), and 10 from wild spotted hyaenas at the Siyafunda Wildlife & Conservation (Limpopo, South Africa).

action unit (AU) wolfdog sun bear wild dog hyaena meerkat lion
AU109 + 110 Nose Wrinkler + Upper Lip Raiser present present present present present present
AU12 Lip Corner Puller present present present present present present
AU16/116 Lower Lip Depressor not coded present not found not found present present
AU25 Lips Parted present present present present present present
AU26 Jaw Drop present present present present present present
AU27 Mouth Stretch present present present present present present