P n	Photosynthetic rate
G s	Stomatal conductance
C i	Intercellular CO2 concentration
T r	Transpiration rate
CE	Carboxylation efficiency
L s	Value of stomatal limitation
WUE	Water use efficiency
A	Leaf area
N mass	Leaf N per unit mass
N area	Leaf N per unit area
PNUE	Photosynthetic nitrogen use efficiency
N mass	Leaf N content per unit of leaf dry mass
M A	Leaf dry mass per unit area
C Chl	Leaf chlorophyll content
V cr	Specific activity of RuBisCO
J mc	Potential rate of photosynthetic electron transport per unit cytochrome f
C B	Ratio of leaf chlorophyll to leaf nitrogen invested in light harvesting
P C	Fractions of leaf N allocated to photosynthetic
P R	Fractions of leaf N allocated to RuBisCO
P B	Fractions of leaf N allocated to bioenergetics
P L	Fractions of leaf N allocated to light-harvesting
NSC	Non-structural carbohydrate
SSRs	The ratio of soluble sugar to starch