(A) Average fold changes (log2FoldChange) of liver, heart, kidney, skin, and brain in 147 metabolites are ranked. Both Sdhd single-hKO (n = 3 mice) and Sdhb/c/d triple-hKO (n = 5 mice) tissues showed the highest increase in succinate, and the highest overlapping decreases in 3-hydroxybutyrate and l-cystine, relative to WT control mice (n = 2). A stringent outlier analysis by ROUT method (Q = 0.1%) detected only 4 outlier metabolites, marked by asterisks. (B) The ratios of succinate to fumarate and oxoglutarate were increased in Sdhb/c/d-hKO mice relative to WT. Data represent mean ± SEM.