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. 2022 Aug 5;16(11):2535–2546. doi: 10.1038/s41396-022-01294-9

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© The Author(s) 2022

Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/.

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Fig. 4 — A Hydrogenase genes distributions assigned by phylum; B genes of associated terminal reductases distributions assigned by phylum. Fermentative hydrogenases (group B, A1 and A2 FeFe-hydrogenases), electron-bifurcating hydrogenases (group A3 and A4 FeFe-hydrogenases), energy-converting hydrogenases (bidirectional; group 4a, 4c, 4d, 4e, 4f and 4g NiFe-hydrogenases), methanogenic hydrogenases (Fe-hydrogenases, group 3a, 3c, 4h, 4i and 1k NiFe-hydrogenases), respiratory hydrogenases (group 1a, 1b, 1c, 1d, 1k, 2a), sensory hydrogenases (group C FeFe-hydrogenases). HydB hydrogenase-associated diaphorase. NifH nitrogenase. H₂ uptake pathways can be coupled to fumarate reduction (FrdA fumarate reductase), nitrate ammonification (NrfA, ammonia-forming nitrite reductase; NarG, dissimilatory nitrate reductase; NapA, periplasmic nitrate reductase), sulfate and sulfite reduction (AprA adenylylsulfate reductase; AsrA alternative sulfite reductase; DsrA, dissimilatory sulfite reductase), dimethyl sulfoxide and trimethylamine N-oxide reduction (DmsA DMSO and TMAO reductase), reductive acetogenesis (AcsB, acetyl-CoA synthase), aerobic respiration (CydA cytochrome bd oxidase), and methanogenesis (McrA methyl-CoM reductase). No rank means there is no specific taxonomic information at the phylum level. Only genes of average relative abundance > 1 were shown, while other were shown in additional file S6. Significance was tested using independent two-group Wilcoxon rank-sum tests. Data with error bars are expressed as mean ± standard error. **p < 0.01, ***p < 0.001, n = 12/group.