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. 2022 Oct 12;13:1000758. doi: 10.3389/fendo.2022.1000758

Table 1.

Characteristics of task-related taVNS studies that are included in the review.

Study(author/year) Sample size Study design & protocol Age(years) Stimulationparameter Symptom Targetdisorder taVNS effects
Colzato et al., 2018 (63) taVNS: 40 Sham: 40
F: 50
Between
Acute taVNS
17-33 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Depression Depressive disorders Divergent thinking
Neuser et al., 2020 (64) 81 (47 F) Within
Acute taVNS
25.3(3.8) taVNS: cymba conchae
1.28(0.58) mA
sham: earlobe
1.82(0.63) mA 25Hz, 30s on 30s off
Depression Depressive disorders Reward seeking
De Smet et al., 2021 (65) taVNS: 42 Sham: 41
F: 66
Between
Acute taVNS
21.11(3.10) taVNS: cymba conchae 1.37(0.81) mA
sham: earlobe
1.89(0.89) mA
25Hz, 250μs,
30s on 30s off
Depression Depressive disorders Negative emotion regulation
Ferstl et al., 2021 (66) 82 (47 F) Within
Acute taVNS
24.6(3.5) taVNS: cymba conchae
sham: earlobe
25Hz, 30s on 30s off
Depression Depressive disorders Mood recovery
Steenbergen et al., 2021 (67) 73 (58 F) Within
Acute taVNS
18-28 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Depression Depressive disorders Recognition of sadness
Koenig et al., 2021 (68) 33 (27 F) adolescents with major depressive disorders Within
Acute taVNS
14-17 taVNS: concha
sham: earlobe
0.5mA, 1Hz, 250μs,
30s on 30s off
Depression Depressive disorders Attention to sad stimuli
Kraus et al., 2007 (38) 6 (5 F) Within
Acute taVNS
20-37 taVNS: inner tragus
sham: earlobe
8Hz, 20μs
Depression Depressive disorders BOLD-signal in limbic brain areas
Subjective well-being
Burger et al.,2020 (69)
(Study 1)
taVNS: 45 Sham: 49
subclinical, high trait worrying sample
Between
Acute taVNS
Not reported taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 250μs,
30s on 30s off
Anxiety and fear GAD Attentional engagement to threat
Burger et al., 2019 (70) taVNS: 48 Sham: 49
subclinical, high trait worrying sample
Between
Acute taVNS
Not reported taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 250μs,
30s ON 30s off
Anxiety and fear GAD
PTSD
Negative thought intrusions
Burger et al., 2017 (71) taVNS: 25 Sham: 26
F: 26
Between
Acute taVNS
20-36 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 250μs,
30s on 30s off
Anxiety and fear Anxiety
PTSD
Extinction of declarative fear
Burger et al., 2016 (72) taVNS: 18 Sham: 13
F: 24
Between
Acute taVNS
18-25 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 250μs,
30s on 30s off
Anxiety and fear Anxiety
PTSD
Extinction learning
Szeska et al., 2020 (73) 80 (57 F)
Fear learning group:
taVNS: 20
sham: 20
control group:
taVNS: 20
sham: 20
Mixed
Acute taVNS
18-34 taVNS: cymba conchae
2.28(1.13) mA
sham: earlobe
2.53(1.11) mA, 25Hz, 200-300μs,
30s on 30s off
Anxiety and fear Anxiety
PTSD
Inhibition of fear potentiated startle responses
Jacobs et al., 2015 (74) 30 (15 F) Within
Acute taVNS
60.57(2.54) taVNS: inner tragus
sham: earlobe
5.0mA, 8Hz, 200μs
Anxiety and fear Anxiety
PTSD
Associated memory performance
Giraudier et al., 2020 (75) 60 (46 F) Between
Acute taVNS
23.45(4.87) taVNS: cymba conchae 1.48(0.59) mA
sham: earlobe
1.31(0.50) mA
25Hz, 200-300μs,
30s on 30s off
Anxiety and fear Anxiety
PTSD
Recollection-based memory
Colzato et al., 2017 (76) 38 (30 F) Within
Acute taVNS
18-26 taVNS: cymba conchae
sham: earlobe
0.5mA,25Hz, 200-300μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Recognition of emotions
Sellaro et al., 2018 (77) 24 (15 F) Within
Acute taVNS
18-28 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Emotion recognition
Zhu et al., 2022 (78) 49 (17 F) Within
Acute taVNS
19.88(1.62) taVNS: tragus
0.86(0.04) mA
sham: earlobe
1.49(0.08) mA
25Hz, 500μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Visual attention towards social salient facial features
Endogenous oxytocin release
Koenig et al., 2021 (68) 30 (24 F) healthy controls Within
Acute taVNS
14-17 taVNS: concha
sham: earlobe
0.5mA, 1Hz, 250μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Emotion recognition
Villani et al., 2019 (79) 46 (32 F) Within
Acute taVNS
21.2(3.1) taVNS: tragus
1.26(0.23) mA
sham: earlobe
1.18(0.18) mA
25Hz, 250μs
Social dysfunction Social dysfunction Interoceptive accuracy
Maraver et al., 2020 (80) 43 (39 F) Within
Acute taVNS
18-30 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Attention to faces with a direct gaze
Steenbergen et al., 2021 (67) 73 (58 F) Within
Acute taVNS
18-28 taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Social dysfunction ASD
ADHD
OCD
Recognition of anger
Ventura-Bort et al., 2021 (81) 37 (20 F) Within
Acute taVNS
23.15 taVNS: cymba conchae
1.34 mA
sham: earlobe
1.58 mA
25Hz, 200-300μs
Social dysfunction ASD
ADHD
OCD
Recollection-based memory for emotional material
Attentional discrimination between emotional and neutral scenes
Steenbergen et al., 2015 (82) 30 (26 F)
taVNS: 15 Sham: 15
Between
Acute taVNS
18-27 taVNS: outer auditory canal
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Responses when two actions were executed in succession
Beste et al., 2016 (83) 51 (37 F)
taVNS: 25 Sham: 26
Between
Acute taVNS
23.63 taVNS: inner ear
sham: earlobe
0.5mA, 25Hz 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
The ability of inhibitory control
Fischer et al., 2018 (84) 21 (18 F) Within
Acute taVNS
20.3(1.4) taVNS: cymba conchae
1.3 mA
sham: earlobe
1.49 mA
25Hz, 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Adaption to conflict
Jongkees et al., 2018 (85) 40 (32 F)
taVNS: 20 Sham: 20
Between
Acute taVNS
taVNS: 22.3(2.7) years Sham: 22.5(2.5) years taVNS: medial of the tragus
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Response selection processes
Keute et al., 2019 (86) 16 (8 F) Within
Acute taVNS
20-28 taVNS: cymba conchae 5.9(1.6) mA
sham: earlobe
7.5(0.8) mA
25Hz, 200μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Automatic motor response inhibition
Keute et al., 2020 (87) 22 (16 F) Within
Acute taVNS
21-28 taVNS: cymba conchae 2.37(0.16) mA
sham: earlobe
2.6 mA
25Hz, 200μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
General adaptive control and sustained attention
Borges et al., 2020 (88) 23 (9 F) Within
Acute taVNS
23.17(4.08) taVNS: cymba conchae
2.19(0.93) mA
sham: earlobe
2.20(1.06) mA 25Hz, 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Cognitive flexibility
Pihlaja et al., 2020 (89) 25 (16 F) Within
Acute taVNS
25.5(4.8) aVNS: inner tragus
sham: earlobe
30Hz, 250μs
Impulsivity and inattention ADHD
ASD
DBD
Cognitive control resources required to withhold a prepotent response
Steenbergen et al., 2020 (90) 84 (52 F) Within
Acute taVNS
22.32(2.71) taVNS: cymba conchae
sham: earlobe
0.5mA, 25Hz, 200-300μs,
30s on 30s off
Impulsivity and inattention ADHD
ASD
DBD
Self-control
Llanos et al. 2020 (91) 36 (20 F)
tVNS-hard: 12
tVNS-easy: 12
control: 12
Between
Acute taVNS
21.60(3.56) taVNS: cymba conchae
hard: 1.67(0.79) mA
easy: 1.24(0.88) mA
control: no stimulation
Others/ Language deficits ASD Speech category learning & retention of correct stimulus response associations
Thakkar et al., 2020 (92) 37 (27 F)
Computer control: 7
Device sham control: 7
Earlobe stimulation control: 9
taVNS: 14
Between
Acute taVNS
18-28 taVNS: cymba conchae 1.68(0.87) mA
sham control: cymba conchae
no stimulation
earlobe control: earlobe
1.51(0.35) mA
5Hz, 200μs,
30s on 30s off
Others/ Language deficits ASD Novel orthography acquisition
Hong et al., 2019 (93) 14 patients requiring open laparotomy (8 F) Within
Acute taVNS
57.6 (10.5) taVNS: cymba conchar
10mA, 25Hz, 250μs,
Control: no stimulation
Others/ Gastrointestinal problems ASD Stomach function
Teckentrup et al., 2020 (94) 22 (14 F) Within
Acute taVNS
19-29 taVNS: cymba conchae 1.37(0.81) mA
sham: earlobe
1.89(0.89) mA
25Hz
30s on 30s off
Others/ Gastrointestinal problems ASD Gastric function
Steidel et al., 2021 (95) HF taVNS: 24 (15 F)
LF taVNS:
28 (18 F)
Mixed
Acute taVNS
25.5(5.2) taVNS: cymba conchae, 250μs HF: 25 Hz 0.91(0.43) mA
LF: 1 Hz
0.66(0.53) mA
30s on 30s off
Others/ Gastrointestinal problems ASD Gastric function
Wu et al., 2021 (96) 40 patients with primary insomnia
Group A: 20 (15 F)
Group B: 20 (13 F)
Between
repeated taVNS (30min of taVNS twice a day, 5 days per week for 4 weeks)
Group A: 49.40±12.22 years
Group B: 46.20±12.76 years
taVNS: cavum concha
7-12mA, 20Hz, 200μs,
No stimulation for Group B (controls)
Others/ Sleep problems GAD
MDD
Sleep quality ↑
Zhang et al., 2021 (42) 20 patients with primary insomnia Within
repeated taVNS (30min of taVNS twice a day, 5 days per week for 4 weeks)
Not reported taVNS: cavum concha
0.8-1.5mA, 4/20 Hz, 200μs,
Others/ Sleep problems GAD
MDD
Score of Pittsburgh Sleep Quality Index
Sleep duration↑
He et al., 2022 (97) 24 patients with chronic insomnia (CI) (12F)
18 healthy controls (HC) (12 F)
Between
repeated taVNS for CI patients (30min of taVNS twice a day for 4 weeks),
CI patients:
42.50±15.42 years
HC:
43.5 ± 11.23
years
taVNS: bilateral cymba conchae 4/20 Hz, 200μs ± 30%
No stimulation for HC
Others/ Sleep problems GAD
MDD
The scores of Pittsburgh Sleep Quality Index and Flinders Fatigue Scale

F, female; HF, high frequency; LF, low frequency. ↑: increased/higher/better. ↓: decreased/lower.