Table 3.

Ethogram for Zoo Monitor. Behaviors recorded whenever they occurred during each 15-min observation period. In addition, the location of each penguin was noted on the map (Figure 1) every 1.5 min during each 15-min observation period.

Behavior	Definition
Allo-preen	One bird uses beak to groom feathers of another
Duet	Pair vocalize together
Courtship	Mount or attempted mount
Keeper enters	Keepers comes into exhibit, usually to feed or enrich
Keeper leaves	Keeper exits exhibit
Aggression	Penguin pecks or chases another