Updated phylogenetic analysis of spinosaurids resolves two stages in the evolution of piscivory and display. We show key cranial adaptations in the skull and highlight changes at the anterior end of the trunk to enhance neck ventroflexion (second dorsal vertebra in lateral and anterior views). Bottom, the fully terrestrial theropod Allosaurus fragilis (Madsen, 1976); middle, the baryonychine spinosaurid Suchomimus tenerensis (MNBH GAD70); top, the spinosaurine Spinosaurus aegyptiacus (BSPG 1912 VIII 19). con, condyle; dr, dental rosette; en, external naris; hy, hypopophysis; k, keel; ncr, nasal crest; ns, neural spine; pa, parapophysis; tp, transverse process.